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Plant propagation
Definition
Plant propagation can also refer to the artificial or natural dispersal of plants. Plant
propagation is the process of creating new plants from a variety of sources: seeds, cuttings,
bulbs and other plant parts. Basically all plants in this universe multiply themselves both by
sexual (seeds) or asexual (vegetative) means. However man has now developed some more
techniques for speedy and better multiplication of these plants.
Plant propagation methods
The first and foremost objective of a propagation method should be to produce individuals
that are identical to mother or original plant. Thus a successful propagation method is one
which transmits all desirable characters of a mother plants to the offspring‟s.
1. Sexual Propagation
The plants which multiply through seeds as a mode of perpetuation. Example: Most annuals,
biennials and many perennial fruit plants; vegetable crops; plantation, aromatic and medicinal
plants; ornamental flowering and shade providing shrubs and trees etc. In general the
selfpollinated plants which are considered homozygous are propagated by seeds. In sexual
method of propagation, the sex organs of flower are involved in process like pollination and
fertilization leads to the formation of seeds. Seeds are typically produced from sexual
reproduction within a species may have different characteristics from its parents. Example:
Papaya, phalsa, mangosteen, rootstocks plants of many fruit crops raised through sexual
method of propagation.
Advantages:
For evolution of new varieties through breeding, the hybrids are raised from seed.
1. In some fruit plants like papaya, this is the most popular method of propagation.
2. Seed propagated rootstocks are hardy and develop better root system.
3. Viruses don„t transmit through seeds, thus mostly the seedlings are free from virus
diseases.
4. Occurrence of polyembryony (more than one embryo in seed) in citrus and mango leads to
the development of uniform seedlings as in asexual method.
Disadvantages
1. Seedlings have a long juvenile period and come into bearing later as compared to asexually
raised plants.
2. Due to segregation of characters, the progeny is not true-to-type.
3. It is not economical to handle larger trees, as less number of trees can be accommodated
per unit area and the cultural operations are difficult.
2. Asexual /Vegetative Propagation
Propagation by apomictic seedlings: Apomictic seedlings are identical to their mother plants,
and similar through the plants raised through other vegetative means, as it has the same
genetic makeup as that of the mother plants. Example: Citrus, mango, apple etc.
Propagation by vegetative structure: It is also called as vegetative propagation as it
involves only vegetative parts without any sexual plant parts. The plant parts like leaf, stem,
root and other root producing plant organs are used. The new individual propagated through
this method is true to type. The commercially important fruit crops are propagated by
vegetative method.
Types of Plant propagation
1. Propagation by Cuttings
A cutting is a piece of vegetative tissue (stem, root or leaf) that, when placed under suitable
environmental conditions, will regenerate the missing parts and produce a self-sustaining
plant.
In this method of propagating fruit plants, a plant (generally stem) having at least few buds,
when detached from parent plant and placed under favourable conditions develop into a
complete plant resembling in all characteristics to the parent from which it was taken.
This method is commonly used in plants, which root easily and readily, thus multiplication of
plants is very quick and cheap.
The fruit plants like Phalsa, pomegranate, lemon and grapes etc. are commercially propagated
by cuttings.
However this method of propagation has certain disadvantages like
 Some plants do not root easily.
 The desired benefit of rootstock of rootstock on a desired variety cannot be exploited.
Cuttings are prepared from vegetative portion of the plants such as stem, root, leaf and are
classified according to the plant part used and are described below:
a. Stem Cutting
A stem cutting is any cutting taken from the main
shoot of a plant or any side shoot growing from the
same plant or stem.
Propagation by stem cuttings is the most commonly used method to propagate many woody
plants. There are several types of stem cuttings.
i) Hardwood cutting
Hardwood cuttings are prepared during dormant
season, usually from one year old immature shoots of
previous season‟s growth.
In case of deciduous plants, the cuttings are made
after pruning.
Generally the cuttings of 15-20 cm length and having
3-5 buds are preferred depending upon species.
While preparing the cutting, a straight cut is given at the base of shoot below the node while a
slanting cut 1-2 cm above the bud is given at the top, e.g. grape, fig, pomegranate, mulberry,
kiwifruit, olive, quince, hazel nut, chest nut, plum, gooseberry and apple.
ii) Semi Hardwood Cutting
Semi hardwood cuttings are prepared from partial matured, slightly woody shoot.
These are succulent and tender in nature and are usually prepared from growing wood of
current season‟s growth.
The length of cutting varies from 7-20 cm.
The cuttings are prepared by trimming the cutting with straight cut below a node and
removing a few lower leaves.
However, it is better to retain two to four leaves on the top of cuttings.
Treating the cutting with 5000 ppm IBA (an rooting hormone) before planting gives better
result.
The best time of taking cutting is summer when new shoots is emerged and their wood is
partially matured.
Ex. Mango, Guava, Jackfruit, Lemon etc.
iii) Soft-Wood Cutting
Softwood cutting is the name given to any cuttings
prepared from soft, succulent and non-lignified
shoots which have not become hard or woody.
Softwood cuttings are taken from new growth of the
current season.
They are still flexible and non-woody. Softwood
cuttings are generally the easiest to root and don‟t
require special handling. e.g. lime and lemon.
b) Leaf bud cutting
There are relatively few plants which can be
reproduced by using a leaf or the portions of a
leaf to produce a new plant.
The reason for this is that leaf cuttings must
regenerate both new root and bud tissues.
Not many plants have this capacity. Most of the
plants propagated by leafcuttings are house
plants with thick, fleshy leaves.
Depending on the plant to be propagated, the
leaf cutting may involve the leaf blade only, or a leaf with petiole, or merely portions of a
leaf.
Leaf-bud cuttings are unlike leaf cuttings in that it contains a portion of stem tissue and most
importantly, a bud.
The bud, located at the junction of the leaf petiole and the stem, is a preformed growing
point.
Leaf cutting should preferably be prepared during growing season because buds if inter in
dormancy may be difficult to force to active stage.
Plant material selected for leaf cuttings should be healthy, actively growing and free of insect
or disease problems.
E.g. Blackberry, lemon, raspberry etc.
c) Root cutting
Root cuttings can be used to propagate a range
of herbaceous perennials in late autumn or
early winter when the plants are dormant. Root
cuttings are used to propagate plants that
naturally produce suckers (new shoots) from
their roots.
Root cuttings require no special aftercare
needed for aerial cuttings as well as large
numbers of new plants can be generated from
each parent plant and the plants derived from root cuttings are relatively large and vigorous.
Another advantage of root cuttings is that plants from root cuttings are free of foliar pests and
pathogens that might affect their parents, such as stem and leaf nematodes.
E.g. blackberry, fig, cherry, raspberry etc.
2. Propagation by Layering
2.1. Air-layering
In this method one-year-old, healthy
and straight shoot is selected and ring of
bark measuring about 2.5 to 4.0 cm just
below a swollen bud is removed.
The cut is then surrounded sphagnum
moss or any other material which can
retain moisture for long period of time,
can be used for this purpose and is
wrapped with a polyethylene strip (200–
400 gauge).
Both ends are tied with fine rope or
rubber bands to make it practically air-
tight and after 30 to 45 days, roots are
formed in the aerial part of the plant.
The ideal time is February–March and
July–August months for air- layering.
Litchi, Lime and sweet lime can be
propagated by air layering.
Application of root promoting hormones at the time of layering helps to get profuse rooting
within a short time.
Root promoting substances may be applied as powder or in lanolin or as a solution.
2.2 Trench layering
In this method, it is important to establish a
permanent row of plants to be propagated.
A branch is laid horizontally in a small trench
to encourage the development of several new
shoots from it.
As these shoots grow, soil is filled around
them and roots eventually develop.
The mother plants are planted at the base of a trench at an angle of 45 0 in rows.
The long and flexible stems of these plants are pegged down on the ground to form a
continuous line of layered plants.
The young shoots then arise from
these plants are gradually
mounded up to a depth of 15-20
cm,
e.g., Apple rootstocks (M16 and
M25), cherry, plum etc.
2.3 Tip layering
In this method of propagation,
bending of a plant stem to the
ground and covering the tip with
soil so that roots and new shoots
may develop.
In tip layering, the tip of shoots
are bend to the ground and the
rooting takes place near the tip of
current season‟s shoot.
2.4 Serpentine/compound layering
Serpentine layering, also known as compound layering is similar to simple layering, except
that multiple sections of the stem are buried, resulting in multiple plants from one stem.
With this technique, a long, vine-like stem is ideal.
It is modification of simple layering in which one year old branch is alternatively covered and
exposed along its length.
e.g. American grape.
2.5 Mound layering /Stooling
In this method, plants are headed back to 15-20 cm above the ground level during dormant
season.
The new shoots come out within two months after heading back.
These sprouts are then girdled near base and rooting hormone (IBA), made in lanolin paste is
applied to the upper portion of cut with moist soil.
The rooting of shoots is observed within 25–35 days.
The stooling is used in the commercial production of clonal rootstocks of temperate fruit
crops like apple, pear etc. and sometimes in guava also.
3. Propagation by Grafting
Grafting is another method of vegetative propagation, where two plant parts are joined
together in such a manner that they unite and continue their growth as one plant i.e. the stock
and scion are placed in close contact with each other and held together firmly, until they unite
to form a composite plant.
The different methods of grafting are tongue grafting, cleft grafting, approach grafting, side
grafting and veneer grafting etc.
3.1. Tongue grafting (Splice/Whip grafting)
It is the simple and popular propagation method
used in apples and widely used in pear.
This method is commonly used when the stock
and scion are of equal diameter.
Each scion sticks should contain at least two to
three sets of buds.
Identical cuts are made at the top of the rootstock and
bottom of the scion, so the two pieces fit together
nicely.
About one-year-old rootstock is headed back at a height
of 23-25 cm from the soil and a diagonal cut is made at
the distal end of the rootstock.
A similar slanting cut is made on the proximal end of
the scion. Try to make this cut with one stroke of the
knife.
The cut surface of both rootstock and scion are bound together and tied firmly. The scion
having 2 to 3 buds is then tightly fitted with the rootstock taking care that the cambium layer
of at least one side of the stock and scion unites together.
This is then wrapped with polyethylene strip. Tongue grafting is done in March–April in high
hills and dry temperate zone while February-March in lower elevation.
Many fruit plant are propagated by
whip grafting.
3. 2. Cleft grafting
This is also known as wedge grafting.
Wedge grafting is a relatively easy
method of propagation.
In this technique, proper selection and
preparation of scion sticks is very
important for obtaining higher
success of graft.
This method is useful in the nursery
where the rootstock is quite thicker
than scion and tongue grafting
cannot be employed successfully.
Rootstock up to 8 cm girth is
selected for this purpose. The
rootstock is cleft grafted after
decapitating the stock 45 cm above
the ground level.
The beheaded rootstock is split to about 5cm deep through the center of stem.
After that a hard wooden wedge is inserted to keep open for the subsequent insertion of scion.
The scion of 15-20 cm size is taken from a terminal shoot, which is more than three month
old and then it is wedge securely (6-7 cm).
The cleft of the scion then slipped into the split of the stock. In thicker rootstock more than
one scion should be inserted.
Care must be exercised to match the cambium layer of the stock and scion along with full
length of each component.
This technology has overcome seasonal barriers and planting materials could be raised
throughout the year either in greenhouse or under open field conditions. Polyethylene cap
facilitates early sprouting and ensures good success rate of grafts, e.g. avocado, apple, pear,
plum, mango etc.
3.3. Side grafting
In this method of grafting, a three
sided rectangular cut about 4.0 x
1.25 cm is given on the rootstock
at a height of about 15-18 cm and
the bark of the demarcated portion
is detached from the rootstock.
A similar cut is also given on the
base of the scion stick to expose
cambium.
The scion should be prepared well
in advance before the actual
grafting.
The healthy scion shoots of previously mature flush are selected. The selected scion shoots
should have plump terminal buds.
After the selection of the scion shoots, remove the leaf blades, leaving petioles attached to the
scion.
In about 8 to 10 days, the attached petioles drop automatically and terminal buds become
swollen.
Now the scion stick should be detached from the mother tree and grafted on the stock.
The prepared scion is inserted under the bark flap of the rootstock after given a slanting cut
so that the exposed cambium of the two components is in close contact with each other.
The bark flap of the rootstock is resorted in its position. The graft union is then tied firmly
with polythene strip.
After the successful completion of the grafting operation, a part of the top of rootstock is
removed to encourage growth of the scion.
When the scion has shown emergence of leaves, the root stock portion above the graft union
should be removed.
Side grafting can be carried out successfully from March to October but success during the
May and October is rather low. This method of propagation is commonly used in mango.
3.4. Inarching or Approach grafting
The method of inarching or
approach grafting is quite
cumbersome and time
consuming, but it is still the
leading method for commercial
propagation of many fruit
plants.
It is generally used for
repairing or replacing damaged root system and hence also called as repair grafting.
This method of grafting is termed approach grafting, as the rootstock is approached to the
scion, while it is still attached to the mother plant.
Selection of parent tree for taking the scion is an important factor for its success.
The scion plant should be healthy, vigorous and high yielding. The stock is brought close to
the scion.
In this method the diameter of rootstock and scion should be approximately the same. A slice
of bark along with a thin piece of wood about 6-8 cm long and about 1/3 inch in thickness at
height is removed from matching portions of both the stock and the scion.
They are then brought together making sure that their cambium layers make contact with
each other.
These grafts are then tied firmly with polythene strip or any other tying material. The stock
and scion plants are watered regularly to hasten the union.
The union is complete in about 2 to 3 months. After successful union, stock above and scion
below the graft union are looped of gradually.
Last week of July or the first week of August is the best period for approach grafting, e.g.
mango, sapota, guava, litchi etc.
3.5. Veneer grafting
This method of propagation possesses promise for mass scale commercial propagation. It is
simple method of propagation.
However, several factors affecting the success like age and diameter of scion, season of
grafting and defoliation period of scion stick etc.
About one year old rootstock is suitable for
this method. However, if the stock attains
suitable thickness (about 1to 1.5 cm dia)
earlier than a year, can be utilized for
rootstock purpose.
Better success is obtained with a scion stick
of 3-6 months of age with lust green leaves
are selected.
The scion sticks are pre-defoliated 5-10 days
prior the grafting leaving the petiole
attached for making the auxiliary and apical
buds active.
Usually, the terminal and next to terminal shoots are most ideal. For conducting this grafting
operation, a downward and inward 25–35 mm long cut is made in the stock at a height of
about 15-20 cm.
At the base of cut, a small shorter cut is given to intersect the first so asto remove the piece of
wood and bark.
The scion stick is given a long slanting cut on one side and a small shortcut on another side to
match the cuts with the rootstock.
Now the scion is inserted in the stock so that the cambium layers come in contact from the
longer side.
The graft union is then tied with polyethylene strip. After the scion remains green for more
than 10 days, the rootstock should be clipped in stages.
When scion growth begins, the shoot of rootstock is removed above the graft union. eg
Mango.
3.6. Softwood grafting
In this case, as the name indicates grafting is done on newly emerged rootstock of 40-60 day-
old.
This is easy and simple
method and can be
practiced throughout the
year raised with 60-90%
success. This method is
useful for in situ grafting
while establishing new
orchards with already
established rootstocks in
the field.
In this method, remove the
leaves of the selected
seedling using sharp knife,
retaining one or two pairs of bottom leaves. Give a transverse cut on the top soft portion of
the seedling.
Make a cleft of 3-4 cm deep in the middle of the decapitated stem of the seedling by giving a
longitudinal cut. A little wood is removed from the inner side of the cleft at the top.
The scion is prepares by mend the cut end of the scion into wedge shape of 3-4cm in length
by chopping off the bark and little wood from the two opposite sides, taking care to retain
some bark on remaining two sides.
Insert the wedge of the scion into the cleft of the seedling taking care that cambium layers of
stock and scion come in perfect contact with each other.
Secure the joint firmly with polythene strip (30 cm long, 2 cm broad and 150 gauge). Cover
the scion with polythene cap (15 cm x 10 cm, 100 gauge) and tie it at the bottom to maintain
humidity and to protect the apical bud from drying.
The cap should not touch the terminal bud. July and August months with high humidity and
moderate temperature are the best for the success of softwood grafting.
This is very famous and successful propagation method in cashew nut, mango, sapota etc.
4. Propagation by Budding
Budding is a method of propagation in which only one bud of desired scion is inserted in the
rootstock.
The ideal condition of budding is when the bark starts slipping both on the stock and scion. It
indicates that the cambium is active.
Important method of budding are shield or tbudding, patch budding, chip budding, ring
budding etc.
4.1. Shield or T- budding
„T‟ budding or shield budding is a special
grafting technique in which the scion
piece is reduced to a single bud.
A small branch with several buds suitable
for T budding on it is often called a bud
stick.
Successful T budding requires that the
scion material have fully-formed, mature,
dormant buds, and that the rootstock be
in a condition of active growth such that
the bark is slipping indicating the
vascular cambium is actively growing,
and the bark can be peeled easily from
the stock piece with little damage.
Bud sticks having plump, healthy buds
are suitable as scions. Leaf blades are
clipped from the budsticks, leaving the
petiole intact.
Budding knives should be kept very sharp, so that as little damage as possible is done to the
bud.
The bud and a small sliver of the wood underneath it are cut from the bud stick using an
upward slicing motion.
The cut should begin about 1/2 to 3/4 inch below the bud, and should go deep enough into the
wood so that when the cut is finished about 1/2 to 3/4 above the bud, the bark and a small
sliver of wood are cut off.
A perpendicular cut across the top of the upward cut will separate it from the bud stick. Buds
must be cut from the bud stick just prior to grafting, otherwise they will dry out.
The bark is carefully slipped from the stem of the rootstock exposing a "pocket" into which
the bud shield can be placed.
Care should be taken not to tear the flaps of bark in the process of spreading them.
If the bark does not slip easily, this indicates that the stock is not in active growth and the
process should be conducted later when active growth has resumed.
The bark flaps are held tightly against the bud as they are wrapped with a budding rubber,
grafting tape or other suitable closure.
This closure must be removed in 2 to 3 weeks after the union has healed. If the material does
not break down, it will girdle the rootstock,
e.g. Citrus, plum, peach, cherry, ber, rose etc.
4.2. Patch budding
A rectangular patch of bark is removed
completely from the stock and replace
with a similar patch of bark containing a
bud of desired variety.
Normally done during the growing season when the bark separates readily from the wood
along the cambial layer. It is successfully used in species having thick bark such as walnut,
pecan nut.
4.3. Chip budding
Chip budding is successful method of budding
when the bark of the stock does not slip easily.
A chip of bark and wood is removed from the
smooth surface between the nodes of the stock.
A chip of similar shape and size is then
removed from the bud wood of desired cultivar.
For which, a 2-3 cm long down ward cut is
made through the bark and slightly in to the wood of the stock.
Then a second cut of about 2.5 cm is made so that it bisects the first cut at an angle of 30-45
0C . in this way the chip of wood is removed from the stock.
The bud chip then slipped in the place of rootstock from where chip has been removed, e.g.
mango, grape etc.
4.4. Ring budding
In ring budding, a complete ring of bark is removed from
the stock and it is completely girdled.
A similar ring of bark containing a bud is removed from
the bud stick and is inserted on to the rootstock. In this
budding both scion and stock should be of same size.
It is utilized in peach, plum, ber, mulberry etc.
5. Tissue Culture/Micropropagation:
The principles of tissue culture can be successfully employed in horticultural crops.
Quite a large number of ornamental plants are reported to respond to propagation by tissue
culture method.
Few such plants are gladiolus, carnation, lily, rose, gerbera, anthurium, magnolia, fern, cacti,
etc. Propagation of ornamental plants by this method is gaining popularity.
Among fruit crops, tissue cultured plants are commercially produced in banana, strawberry
etc..
The supply of tissue cultured plants of Grand Nain variety of banana has revolutionised the
banana industry.
Advantages of vegetative propagation
 The horticultural crops which do not produce viable seeds are propagated by vegetative
method.
 Most of the important fruit crops are cross pollinated and are highly heterozygous. When
propagated through seeds, the progenies shows large variation, so vegetative propagation is
remedy for these crops.
 The asexual propagation method gives true to type plants.
 The vegetative way propagated plants bear fruits early.
 In case of fruit crops where root stocks are used, the root stocks impart insect or disease
resistance to the plant.
 Vegetative propagation helps to alter the size of the plant. i.e. dwarfing effect. This helps
for spraying, intercropping & harvesting of crops easy and economical.
 By grafting method different variety of fruit crop can be grown & harvested.
 Inferior quality fruit plants can be converted into good quality plants.
 By means of bridge grafting a repairing of injured plants can be done.
Disadvantages of the vegetative propagation:
 By vegetative propagation new variety cannot be developed.
 It is an expensive method of propagation and required specialized skill.
 The life span of vegetatively propagated plants is short as compared to sexually propagated
plants.
 As all the plants are homozygous the whole plantation may get attacked by a particular pest
or disease.
 Viral diseases could be transferred through vegetative parts.
Recommended Propagation Techniques for Fruit Crops
Fruit crop Commercial propagation method
Almond T-budding and wedge grafting
Aonla Patch budding and wedge grafting
Apple T-budding/tongue and wedge grafting
Apricot T-budding and wedge grafting
Avocado T-budding and wedge grafting
Bael Wedge grafting
Banana Suckers/corm
Cashew Soft wood grafting
Cherry Tongue and wedge grafting
Custard apple Wedge grafting
Date palm Sucker/off shoot
Fig Hard wood/semi-hard wood cutting
Grape Hard wood cutting and wedge grafting
Guava Wedge grafting
Gooseberry Hardwood/semi-hard wood cutting
Jackfruit Patch budding and soft wood grafting
Jamun Soft wood grafting
Kiwi fruit Hard/semi hard wood cutting
Lemon/lime Cutting and budding
Litchi Air layering and wedge
Mandarin T-budding and wedge grafting
Mango Soft wood, wedge and veneer grafting
Peach T-budding, wedge and tongue grafting
Pear T-budding, wedge and tongue grafting
Pecan nut Patch budding and wedge grafting
Pineapple Slip/sucker
Plum T-budding, tongue and wedge grafting
Pomegranate Wedge grafting/air layering, Hard wood
cutting
Raspberry, Blackberry Sucker
Sapota Wedge grafting
Strawberry Runner
Sweet Orange T-budding/wedge grafting
Walnut Walnut Patch budding and wedge grafting
REFERENCE: A.K. Goswami, R.R. Sharma, M. Srivastav, A. Nagaraja and Jai
Prakash. Mthod of propogation in fruit crops. Biotecharticles.
https://www.biotecharticles.com
Dormancyis a period in
an organism's life cycle when growth,
development, and (in animals) physical
activity are temporarily stopped. This
minimizes metabolic activity and
therefore helps an organism to
conserve energy.
Seed dormancy can be defined as the
state or a condition in which seeds are
prevented from germinating even under
the favourable environmental
conditions for germination including,
temperature, water, light, gas, seed
coats, and other mechanical
restrictions.
The main reason behind these
conditions is that they require a period of rest before being capable of germination. These
conditions may vary from days to months and even years. These conditions are the
combination of light, water, heat, gases, seed coats and hormone structures.
Breaking of seed dormancy
Several techniques have been developed to break seed dormancy of different
types. Some of them are:
1. Scarification: Dormancy caused due to hard or impermeable seed coat can be broken by
scarification of the seed coat. Scarification can be done mechanically either by rubbing the seed
with sand paper or by help of scarifier. Chemical scarification is done by treating seeds with
concentrated HCl or H2SO4 for a specified time followed by thorough washing. e.g. pulses,
cotton, seeds of forest trees, etc.
2. Stratification: Dormancy in seeds of many woody and herbaceous species is broken
when hydrated seeds are exposed to relatively low temperatures (00
to 200
C) for 3 to 7 days.
3. High temperature treatment: Dry heating of seeds at 400
–550
C temperature for several
days have been found effective in breaking dormancy in paddy, groundnut, etc.
4. Alternating temperature treatment: Alternating low and high temperature treatment on
imbibed seeds has been found effective for breaking dormancy in many species. e.g. brinjal,
Malus sylvestris, etc.
5. Washing: Washing of dormant seeds by water has been reported to break dormancy,
probably due to leaching of dormancy factor from the seeds. e.g. ragi, groundnut, etc.
6. Dry storage: The embryo of seeds of certain species is not fully developed even at
harvestable maturity stage and requires some more time after harvest (after ripening period)
for full development of embryo. Dry storage of such seeds for few days will break dormancy
and promote germination.
7. Chemical treatments: Various chemicals and growth hormones, viz. Gibberellins,
cytokinins, ethylene, KNO3, thiourea, HNO3, cyanide, etc. have been found to break seed
dormancy in different species. The concentration of the chemicals and mode of treatment
varies in seeds.
Polymorphism of dormancy: In sunflower, the seeds formed in the peripheral portion of the
head/capitulum may not be dormant while those formed in the central portion are dormant.
This is known as polymorphism in seed dormancy.
Classification of fruit crops
Classification is the system of grouping or placing of individuals according to nomenclature.
It is very useful to the pomologist. It helps to:
 To identify and naming the crop.
 To study the close relationship.
 To know their hybrids and crossing behaviour.
 To know their compatibility & inter grafting ability.
 To know their adoptability to soil & climate.
POMOLOGY:
 Pomology is a branch of horticulture which deals with study of various aspects of
fruits loke, rising of saplings, growing them properly and providing various
intercultural operations.
 The term pomology is a combination of two Latin words „Pome‟ means ‗Fruits‟ and
‗Logos„ means ‗study‟.
 “Poma” in Greek also meaning fruits later subsequently “Pome” in Latin word means
fruits, logosstudy.
 Basic Pomology: Study of basic aspects of fruit production like training, water
management, use of PGR„s.
 Commercial Pomology: It is concerned with commercial production of fruits.
 Systematic Pomology: It may be concerned with classification and nomenclature like
kingdom, order, class, genus and species.
A. Classification of fruits based on climate adaptability.
In this classification, the fruits trees are categorized into three recognized groups.
i. Temperate fruits:
 Temperate fruit plants are exacting in their climate requirement.
 They are grown only in place where winter is distinctly cold, require as exposure of specific
chilling temperature for certain period without which they do not flower.
 These fruit plants are generally deciduous and stand frost. Eg. Apple, almond, peach, pear,
plum, strawberry, apricot, persimmon, cherymoya, walnut, peanut, hassle nut, cherry,
pistachios and kiwifruits etc.
ii. Tropical fruits:
 Tropical fruit plants are generally evergreen and are extremely sensitive to cold.
 They do well under lesser fluctuations of diurnal temperature, light and dark periods they
require a moist warm climate but are capable of withstanding dry weather in some cases Eg;
mango, banana, papaya, sapota, etc.,
iii. Sub-tropical fruits:
 The fruit crops grown under a climatic condition between temperate and the tropical are
known as subtropical fruit crops.
They may be either deciduous or evergreen and are usually able to withstand a low
temperature but not the frost.
 They are also quite adoptive to fluctuations of light and dark period during day and night.
 Some subtropical fruit plants require chilling for flower bud differentiation.
 Example; grape, citrus, durian, jackfruit, etc.,
B. Classification based on bearing habit:
On the basis of bearing habit, fruit trees are classified in to six categories to facilitate cultural
operation like pruning, skiffing, heading back etc.
1. Fruit buds bore terminally and giving rise to inflorescence without leaves e.g. Mango,
Cherry, etc.
2. Fruit buds borne terminally and unfolding to produce leafy shoots which terminate in
flower clusters. e.g. Apple
3. Fruit buds borne terminally and unfolding to produce leafy shoots with flower or flower
clusters e.g Guava
4. Fruit bud borne laterally containing flower parts only and giving rise to inflorescence
without leaves or leaves present, they are reduced in size., e.g. Citrus
5. Fruit bud borne laterally and unfolding to produce leafy shoots terminally in flower
clusters this type of flowering is noticed in grapes and cashewnut.
6. Fruit buds borne laterally and unfolding to produce leafy shoots with flower clusters in
leafy axils. eg.Fig.
C. Fruit morphology:
1) Simple fruit - Berry: Banana, Papaya, Grape, Sapota, and Avocado
2) Modified berry
i. Balusta : Pomegranate
ii. Amphisarca : Woodapple, Bael
iii. Pepo : Water melon
iv. Pome : Apple, Pear, Laquat
v. Drupe (Stone) : Mango, Pear, Plum
vi. Hesperidium : Citrus
vii. Nut : Cashew, Litchi, Walnut, Rambutan
viii. Capsule : Anola, Carambola
3) Aggregate fruits : Etario of berries –Custard apple, Raspberry
4) Multiple fruit : Syconus- Fig : Sorosis- Jackfruit, Pineapple, Mulberry
D. Based on rate of respiration:
Climacteric Fruits Non-climacteric Fruits
Mango, Banana, Sapota, Guava, Papaya,
Apple, Fig, Peach, Pear, Plum, Annona,
Tomato
Citrus, Grape, Pomegranate Pineapple Litchi,
Ber, Jamun, Cashew, Cucumber, Cherry,
Strawberry
(Climacteric fruits produce much larger amount of ethylene than non climacteric fruits)
E. Based on photoperiodic responses
Long day Short day Day neutral plant
Passionfruit, Banana, Apple Strawberry, Pineapple,
Coffee
Papaya, Guava
F. Based on relative salt tolerance
Highly tolerant Medium tolerant Highly sensitive
Datepalm, Ber, Amla, Guava,
Coconut, Khirni
Pomegranate, Cashew, Fig,
Jamun, Phalsa
Mango, Apple, Citrus, Pear,
Straw berry
G. Based on relative acid Tolerance
Highly tolerant Medium tolerant Highly sensitive
Stawberry, Raspberry, Fig,
Bael, Plum
Pineapple, Avocado, Litchi
H. Based on longevity:
a) Very Long longevity - >100yrs- Datepalm, Coconut, Arecanut
b) Long longevity - 50-100yrs - Mango, Tamarind
c) Medium - 10-50yrs - Litchi, Guava,Pomogranate
d) Short - Pineapple, Banana
I. Based on consumers preference or weight of fruits
a) Very light - 50-100gm Grape,Ber,Banana
b) Light - 100-150gm Sapota,Pomegranate
c) Light medium - 150-300gm Mango
d) Medium - 300-350gm Avocado
e) Medium to heavy - 800-1000gm Mango
f) Heavy - 1-5kg Bread fruit, Pineapple
g) Very heavy - >5kg Jack Fruit
Area, production, productivity and export potential

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Plant propagation

  • 1. Plant propagation Definition Plant propagation can also refer to the artificial or natural dispersal of plants. Plant propagation is the process of creating new plants from a variety of sources: seeds, cuttings, bulbs and other plant parts. Basically all plants in this universe multiply themselves both by sexual (seeds) or asexual (vegetative) means. However man has now developed some more techniques for speedy and better multiplication of these plants. Plant propagation methods The first and foremost objective of a propagation method should be to produce individuals that are identical to mother or original plant. Thus a successful propagation method is one which transmits all desirable characters of a mother plants to the offspring‟s. 1. Sexual Propagation The plants which multiply through seeds as a mode of perpetuation. Example: Most annuals, biennials and many perennial fruit plants; vegetable crops; plantation, aromatic and medicinal plants; ornamental flowering and shade providing shrubs and trees etc. In general the selfpollinated plants which are considered homozygous are propagated by seeds. In sexual method of propagation, the sex organs of flower are involved in process like pollination and fertilization leads to the formation of seeds. Seeds are typically produced from sexual reproduction within a species may have different characteristics from its parents. Example: Papaya, phalsa, mangosteen, rootstocks plants of many fruit crops raised through sexual method of propagation. Advantages: For evolution of new varieties through breeding, the hybrids are raised from seed. 1. In some fruit plants like papaya, this is the most popular method of propagation. 2. Seed propagated rootstocks are hardy and develop better root system. 3. Viruses don„t transmit through seeds, thus mostly the seedlings are free from virus diseases. 4. Occurrence of polyembryony (more than one embryo in seed) in citrus and mango leads to the development of uniform seedlings as in asexual method. Disadvantages 1. Seedlings have a long juvenile period and come into bearing later as compared to asexually raised plants. 2. Due to segregation of characters, the progeny is not true-to-type.
  • 2. 3. It is not economical to handle larger trees, as less number of trees can be accommodated per unit area and the cultural operations are difficult. 2. Asexual /Vegetative Propagation Propagation by apomictic seedlings: Apomictic seedlings are identical to their mother plants, and similar through the plants raised through other vegetative means, as it has the same genetic makeup as that of the mother plants. Example: Citrus, mango, apple etc. Propagation by vegetative structure: It is also called as vegetative propagation as it involves only vegetative parts without any sexual plant parts. The plant parts like leaf, stem, root and other root producing plant organs are used. The new individual propagated through this method is true to type. The commercially important fruit crops are propagated by vegetative method. Types of Plant propagation 1. Propagation by Cuttings A cutting is a piece of vegetative tissue (stem, root or leaf) that, when placed under suitable environmental conditions, will regenerate the missing parts and produce a self-sustaining plant. In this method of propagating fruit plants, a plant (generally stem) having at least few buds, when detached from parent plant and placed under favourable conditions develop into a complete plant resembling in all characteristics to the parent from which it was taken. This method is commonly used in plants, which root easily and readily, thus multiplication of plants is very quick and cheap. The fruit plants like Phalsa, pomegranate, lemon and grapes etc. are commercially propagated by cuttings. However this method of propagation has certain disadvantages like  Some plants do not root easily.  The desired benefit of rootstock of rootstock on a desired variety cannot be exploited. Cuttings are prepared from vegetative portion of the plants such as stem, root, leaf and are classified according to the plant part used and are described below: a. Stem Cutting A stem cutting is any cutting taken from the main shoot of a plant or any side shoot growing from the same plant or stem.
  • 3. Propagation by stem cuttings is the most commonly used method to propagate many woody plants. There are several types of stem cuttings. i) Hardwood cutting Hardwood cuttings are prepared during dormant season, usually from one year old immature shoots of previous season‟s growth. In case of deciduous plants, the cuttings are made after pruning. Generally the cuttings of 15-20 cm length and having 3-5 buds are preferred depending upon species. While preparing the cutting, a straight cut is given at the base of shoot below the node while a slanting cut 1-2 cm above the bud is given at the top, e.g. grape, fig, pomegranate, mulberry, kiwifruit, olive, quince, hazel nut, chest nut, plum, gooseberry and apple. ii) Semi Hardwood Cutting Semi hardwood cuttings are prepared from partial matured, slightly woody shoot. These are succulent and tender in nature and are usually prepared from growing wood of current season‟s growth. The length of cutting varies from 7-20 cm. The cuttings are prepared by trimming the cutting with straight cut below a node and removing a few lower leaves. However, it is better to retain two to four leaves on the top of cuttings. Treating the cutting with 5000 ppm IBA (an rooting hormone) before planting gives better result. The best time of taking cutting is summer when new shoots is emerged and their wood is partially matured. Ex. Mango, Guava, Jackfruit, Lemon etc.
  • 4. iii) Soft-Wood Cutting Softwood cutting is the name given to any cuttings prepared from soft, succulent and non-lignified shoots which have not become hard or woody. Softwood cuttings are taken from new growth of the current season. They are still flexible and non-woody. Softwood cuttings are generally the easiest to root and don‟t require special handling. e.g. lime and lemon. b) Leaf bud cutting There are relatively few plants which can be reproduced by using a leaf or the portions of a leaf to produce a new plant. The reason for this is that leaf cuttings must regenerate both new root and bud tissues. Not many plants have this capacity. Most of the plants propagated by leafcuttings are house plants with thick, fleshy leaves. Depending on the plant to be propagated, the leaf cutting may involve the leaf blade only, or a leaf with petiole, or merely portions of a leaf. Leaf-bud cuttings are unlike leaf cuttings in that it contains a portion of stem tissue and most importantly, a bud. The bud, located at the junction of the leaf petiole and the stem, is a preformed growing point. Leaf cutting should preferably be prepared during growing season because buds if inter in dormancy may be difficult to force to active stage. Plant material selected for leaf cuttings should be healthy, actively growing and free of insect or disease problems. E.g. Blackberry, lemon, raspberry etc.
  • 5. c) Root cutting Root cuttings can be used to propagate a range of herbaceous perennials in late autumn or early winter when the plants are dormant. Root cuttings are used to propagate plants that naturally produce suckers (new shoots) from their roots. Root cuttings require no special aftercare needed for aerial cuttings as well as large numbers of new plants can be generated from each parent plant and the plants derived from root cuttings are relatively large and vigorous. Another advantage of root cuttings is that plants from root cuttings are free of foliar pests and pathogens that might affect their parents, such as stem and leaf nematodes. E.g. blackberry, fig, cherry, raspberry etc. 2. Propagation by Layering 2.1. Air-layering In this method one-year-old, healthy and straight shoot is selected and ring of bark measuring about 2.5 to 4.0 cm just below a swollen bud is removed. The cut is then surrounded sphagnum moss or any other material which can retain moisture for long period of time, can be used for this purpose and is wrapped with a polyethylene strip (200– 400 gauge). Both ends are tied with fine rope or rubber bands to make it practically air- tight and after 30 to 45 days, roots are formed in the aerial part of the plant. The ideal time is February–March and July–August months for air- layering. Litchi, Lime and sweet lime can be propagated by air layering.
  • 6. Application of root promoting hormones at the time of layering helps to get profuse rooting within a short time. Root promoting substances may be applied as powder or in lanolin or as a solution. 2.2 Trench layering In this method, it is important to establish a permanent row of plants to be propagated. A branch is laid horizontally in a small trench to encourage the development of several new shoots from it. As these shoots grow, soil is filled around them and roots eventually develop. The mother plants are planted at the base of a trench at an angle of 45 0 in rows. The long and flexible stems of these plants are pegged down on the ground to form a continuous line of layered plants. The young shoots then arise from these plants are gradually mounded up to a depth of 15-20 cm, e.g., Apple rootstocks (M16 and M25), cherry, plum etc. 2.3 Tip layering In this method of propagation, bending of a plant stem to the ground and covering the tip with soil so that roots and new shoots may develop. In tip layering, the tip of shoots are bend to the ground and the rooting takes place near the tip of current season‟s shoot.
  • 7. 2.4 Serpentine/compound layering Serpentine layering, also known as compound layering is similar to simple layering, except that multiple sections of the stem are buried, resulting in multiple plants from one stem. With this technique, a long, vine-like stem is ideal. It is modification of simple layering in which one year old branch is alternatively covered and exposed along its length. e.g. American grape. 2.5 Mound layering /Stooling In this method, plants are headed back to 15-20 cm above the ground level during dormant season. The new shoots come out within two months after heading back. These sprouts are then girdled near base and rooting hormone (IBA), made in lanolin paste is applied to the upper portion of cut with moist soil. The rooting of shoots is observed within 25–35 days. The stooling is used in the commercial production of clonal rootstocks of temperate fruit crops like apple, pear etc. and sometimes in guava also. 3. Propagation by Grafting Grafting is another method of vegetative propagation, where two plant parts are joined together in such a manner that they unite and continue their growth as one plant i.e. the stock and scion are placed in close contact with each other and held together firmly, until they unite to form a composite plant. The different methods of grafting are tongue grafting, cleft grafting, approach grafting, side grafting and veneer grafting etc. 3.1. Tongue grafting (Splice/Whip grafting) It is the simple and popular propagation method used in apples and widely used in pear. This method is commonly used when the stock and scion are of equal diameter. Each scion sticks should contain at least two to three sets of buds.
  • 8. Identical cuts are made at the top of the rootstock and bottom of the scion, so the two pieces fit together nicely. About one-year-old rootstock is headed back at a height of 23-25 cm from the soil and a diagonal cut is made at the distal end of the rootstock. A similar slanting cut is made on the proximal end of the scion. Try to make this cut with one stroke of the knife. The cut surface of both rootstock and scion are bound together and tied firmly. The scion having 2 to 3 buds is then tightly fitted with the rootstock taking care that the cambium layer of at least one side of the stock and scion unites together. This is then wrapped with polyethylene strip. Tongue grafting is done in March–April in high hills and dry temperate zone while February-March in lower elevation. Many fruit plant are propagated by whip grafting. 3. 2. Cleft grafting This is also known as wedge grafting. Wedge grafting is a relatively easy method of propagation. In this technique, proper selection and preparation of scion sticks is very important for obtaining higher success of graft. This method is useful in the nursery where the rootstock is quite thicker than scion and tongue grafting cannot be employed successfully. Rootstock up to 8 cm girth is selected for this purpose. The rootstock is cleft grafted after decapitating the stock 45 cm above the ground level.
  • 9. The beheaded rootstock is split to about 5cm deep through the center of stem. After that a hard wooden wedge is inserted to keep open for the subsequent insertion of scion. The scion of 15-20 cm size is taken from a terminal shoot, which is more than three month old and then it is wedge securely (6-7 cm). The cleft of the scion then slipped into the split of the stock. In thicker rootstock more than one scion should be inserted. Care must be exercised to match the cambium layer of the stock and scion along with full length of each component. This technology has overcome seasonal barriers and planting materials could be raised throughout the year either in greenhouse or under open field conditions. Polyethylene cap facilitates early sprouting and ensures good success rate of grafts, e.g. avocado, apple, pear, plum, mango etc. 3.3. Side grafting In this method of grafting, a three sided rectangular cut about 4.0 x 1.25 cm is given on the rootstock at a height of about 15-18 cm and the bark of the demarcated portion is detached from the rootstock. A similar cut is also given on the base of the scion stick to expose cambium. The scion should be prepared well in advance before the actual grafting. The healthy scion shoots of previously mature flush are selected. The selected scion shoots should have plump terminal buds. After the selection of the scion shoots, remove the leaf blades, leaving petioles attached to the scion. In about 8 to 10 days, the attached petioles drop automatically and terminal buds become swollen. Now the scion stick should be detached from the mother tree and grafted on the stock.
  • 10. The prepared scion is inserted under the bark flap of the rootstock after given a slanting cut so that the exposed cambium of the two components is in close contact with each other. The bark flap of the rootstock is resorted in its position. The graft union is then tied firmly with polythene strip. After the successful completion of the grafting operation, a part of the top of rootstock is removed to encourage growth of the scion. When the scion has shown emergence of leaves, the root stock portion above the graft union should be removed. Side grafting can be carried out successfully from March to October but success during the May and October is rather low. This method of propagation is commonly used in mango. 3.4. Inarching or Approach grafting The method of inarching or approach grafting is quite cumbersome and time consuming, but it is still the leading method for commercial propagation of many fruit plants. It is generally used for repairing or replacing damaged root system and hence also called as repair grafting. This method of grafting is termed approach grafting, as the rootstock is approached to the scion, while it is still attached to the mother plant. Selection of parent tree for taking the scion is an important factor for its success. The scion plant should be healthy, vigorous and high yielding. The stock is brought close to the scion.
  • 11. In this method the diameter of rootstock and scion should be approximately the same. A slice of bark along with a thin piece of wood about 6-8 cm long and about 1/3 inch in thickness at height is removed from matching portions of both the stock and the scion. They are then brought together making sure that their cambium layers make contact with each other. These grafts are then tied firmly with polythene strip or any other tying material. The stock and scion plants are watered regularly to hasten the union. The union is complete in about 2 to 3 months. After successful union, stock above and scion below the graft union are looped of gradually. Last week of July or the first week of August is the best period for approach grafting, e.g. mango, sapota, guava, litchi etc. 3.5. Veneer grafting This method of propagation possesses promise for mass scale commercial propagation. It is simple method of propagation. However, several factors affecting the success like age and diameter of scion, season of grafting and defoliation period of scion stick etc. About one year old rootstock is suitable for this method. However, if the stock attains suitable thickness (about 1to 1.5 cm dia) earlier than a year, can be utilized for rootstock purpose. Better success is obtained with a scion stick of 3-6 months of age with lust green leaves are selected. The scion sticks are pre-defoliated 5-10 days prior the grafting leaving the petiole attached for making the auxiliary and apical buds active. Usually, the terminal and next to terminal shoots are most ideal. For conducting this grafting operation, a downward and inward 25–35 mm long cut is made in the stock at a height of about 15-20 cm. At the base of cut, a small shorter cut is given to intersect the first so asto remove the piece of wood and bark.
  • 12. The scion stick is given a long slanting cut on one side and a small shortcut on another side to match the cuts with the rootstock. Now the scion is inserted in the stock so that the cambium layers come in contact from the longer side. The graft union is then tied with polyethylene strip. After the scion remains green for more than 10 days, the rootstock should be clipped in stages. When scion growth begins, the shoot of rootstock is removed above the graft union. eg Mango. 3.6. Softwood grafting In this case, as the name indicates grafting is done on newly emerged rootstock of 40-60 day- old. This is easy and simple method and can be practiced throughout the year raised with 60-90% success. This method is useful for in situ grafting while establishing new orchards with already established rootstocks in the field. In this method, remove the leaves of the selected seedling using sharp knife, retaining one or two pairs of bottom leaves. Give a transverse cut on the top soft portion of the seedling. Make a cleft of 3-4 cm deep in the middle of the decapitated stem of the seedling by giving a longitudinal cut. A little wood is removed from the inner side of the cleft at the top. The scion is prepares by mend the cut end of the scion into wedge shape of 3-4cm in length by chopping off the bark and little wood from the two opposite sides, taking care to retain some bark on remaining two sides. Insert the wedge of the scion into the cleft of the seedling taking care that cambium layers of stock and scion come in perfect contact with each other.
  • 13. Secure the joint firmly with polythene strip (30 cm long, 2 cm broad and 150 gauge). Cover the scion with polythene cap (15 cm x 10 cm, 100 gauge) and tie it at the bottom to maintain humidity and to protect the apical bud from drying. The cap should not touch the terminal bud. July and August months with high humidity and moderate temperature are the best for the success of softwood grafting. This is very famous and successful propagation method in cashew nut, mango, sapota etc. 4. Propagation by Budding Budding is a method of propagation in which only one bud of desired scion is inserted in the rootstock. The ideal condition of budding is when the bark starts slipping both on the stock and scion. It indicates that the cambium is active. Important method of budding are shield or tbudding, patch budding, chip budding, ring budding etc. 4.1. Shield or T- budding „T‟ budding or shield budding is a special grafting technique in which the scion piece is reduced to a single bud. A small branch with several buds suitable for T budding on it is often called a bud stick. Successful T budding requires that the scion material have fully-formed, mature, dormant buds, and that the rootstock be in a condition of active growth such that the bark is slipping indicating the vascular cambium is actively growing, and the bark can be peeled easily from the stock piece with little damage. Bud sticks having plump, healthy buds are suitable as scions. Leaf blades are clipped from the budsticks, leaving the petiole intact. Budding knives should be kept very sharp, so that as little damage as possible is done to the bud.
  • 14. The bud and a small sliver of the wood underneath it are cut from the bud stick using an upward slicing motion. The cut should begin about 1/2 to 3/4 inch below the bud, and should go deep enough into the wood so that when the cut is finished about 1/2 to 3/4 above the bud, the bark and a small sliver of wood are cut off. A perpendicular cut across the top of the upward cut will separate it from the bud stick. Buds must be cut from the bud stick just prior to grafting, otherwise they will dry out. The bark is carefully slipped from the stem of the rootstock exposing a "pocket" into which the bud shield can be placed. Care should be taken not to tear the flaps of bark in the process of spreading them. If the bark does not slip easily, this indicates that the stock is not in active growth and the process should be conducted later when active growth has resumed. The bark flaps are held tightly against the bud as they are wrapped with a budding rubber, grafting tape or other suitable closure. This closure must be removed in 2 to 3 weeks after the union has healed. If the material does not break down, it will girdle the rootstock, e.g. Citrus, plum, peach, cherry, ber, rose etc. 4.2. Patch budding A rectangular patch of bark is removed completely from the stock and replace with a similar patch of bark containing a bud of desired variety. Normally done during the growing season when the bark separates readily from the wood along the cambial layer. It is successfully used in species having thick bark such as walnut, pecan nut. 4.3. Chip budding Chip budding is successful method of budding when the bark of the stock does not slip easily. A chip of bark and wood is removed from the smooth surface between the nodes of the stock. A chip of similar shape and size is then removed from the bud wood of desired cultivar. For which, a 2-3 cm long down ward cut is
  • 15. made through the bark and slightly in to the wood of the stock. Then a second cut of about 2.5 cm is made so that it bisects the first cut at an angle of 30-45 0C . in this way the chip of wood is removed from the stock. The bud chip then slipped in the place of rootstock from where chip has been removed, e.g. mango, grape etc. 4.4. Ring budding In ring budding, a complete ring of bark is removed from the stock and it is completely girdled. A similar ring of bark containing a bud is removed from the bud stick and is inserted on to the rootstock. In this budding both scion and stock should be of same size. It is utilized in peach, plum, ber, mulberry etc. 5. Tissue Culture/Micropropagation: The principles of tissue culture can be successfully employed in horticultural crops. Quite a large number of ornamental plants are reported to respond to propagation by tissue culture method. Few such plants are gladiolus, carnation, lily, rose, gerbera, anthurium, magnolia, fern, cacti, etc. Propagation of ornamental plants by this method is gaining popularity. Among fruit crops, tissue cultured plants are commercially produced in banana, strawberry etc.. The supply of tissue cultured plants of Grand Nain variety of banana has revolutionised the banana industry. Advantages of vegetative propagation  The horticultural crops which do not produce viable seeds are propagated by vegetative method.  Most of the important fruit crops are cross pollinated and are highly heterozygous. When propagated through seeds, the progenies shows large variation, so vegetative propagation is remedy for these crops.  The asexual propagation method gives true to type plants.  The vegetative way propagated plants bear fruits early.
  • 16.  In case of fruit crops where root stocks are used, the root stocks impart insect or disease resistance to the plant.  Vegetative propagation helps to alter the size of the plant. i.e. dwarfing effect. This helps for spraying, intercropping & harvesting of crops easy and economical.  By grafting method different variety of fruit crop can be grown & harvested.  Inferior quality fruit plants can be converted into good quality plants.  By means of bridge grafting a repairing of injured plants can be done. Disadvantages of the vegetative propagation:  By vegetative propagation new variety cannot be developed.  It is an expensive method of propagation and required specialized skill.  The life span of vegetatively propagated plants is short as compared to sexually propagated plants.  As all the plants are homozygous the whole plantation may get attacked by a particular pest or disease.  Viral diseases could be transferred through vegetative parts. Recommended Propagation Techniques for Fruit Crops Fruit crop Commercial propagation method Almond T-budding and wedge grafting Aonla Patch budding and wedge grafting Apple T-budding/tongue and wedge grafting Apricot T-budding and wedge grafting Avocado T-budding and wedge grafting Bael Wedge grafting Banana Suckers/corm Cashew Soft wood grafting Cherry Tongue and wedge grafting
  • 17. Custard apple Wedge grafting Date palm Sucker/off shoot Fig Hard wood/semi-hard wood cutting Grape Hard wood cutting and wedge grafting Guava Wedge grafting Gooseberry Hardwood/semi-hard wood cutting Jackfruit Patch budding and soft wood grafting Jamun Soft wood grafting Kiwi fruit Hard/semi hard wood cutting Lemon/lime Cutting and budding Litchi Air layering and wedge Mandarin T-budding and wedge grafting Mango Soft wood, wedge and veneer grafting Peach T-budding, wedge and tongue grafting Pear T-budding, wedge and tongue grafting Pecan nut Patch budding and wedge grafting Pineapple Slip/sucker Plum T-budding, tongue and wedge grafting Pomegranate Wedge grafting/air layering, Hard wood cutting Raspberry, Blackberry Sucker Sapota Wedge grafting Strawberry Runner Sweet Orange T-budding/wedge grafting Walnut Walnut Patch budding and wedge grafting
  • 18. REFERENCE: A.K. Goswami, R.R. Sharma, M. Srivastav, A. Nagaraja and Jai Prakash. Mthod of propogation in fruit crops. Biotecharticles. https://www.biotecharticles.com
  • 19. Dormancyis a period in an organism's life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolic activity and therefore helps an organism to conserve energy. Seed dormancy can be defined as the state or a condition in which seeds are prevented from germinating even under the favourable environmental conditions for germination including, temperature, water, light, gas, seed coats, and other mechanical restrictions. The main reason behind these conditions is that they require a period of rest before being capable of germination. These conditions may vary from days to months and even years. These conditions are the combination of light, water, heat, gases, seed coats and hormone structures. Breaking of seed dormancy Several techniques have been developed to break seed dormancy of different types. Some of them are: 1. Scarification: Dormancy caused due to hard or impermeable seed coat can be broken by scarification of the seed coat. Scarification can be done mechanically either by rubbing the seed with sand paper or by help of scarifier. Chemical scarification is done by treating seeds with concentrated HCl or H2SO4 for a specified time followed by thorough washing. e.g. pulses, cotton, seeds of forest trees, etc. 2. Stratification: Dormancy in seeds of many woody and herbaceous species is broken when hydrated seeds are exposed to relatively low temperatures (00 to 200 C) for 3 to 7 days. 3. High temperature treatment: Dry heating of seeds at 400 –550 C temperature for several days have been found effective in breaking dormancy in paddy, groundnut, etc. 4. Alternating temperature treatment: Alternating low and high temperature treatment on imbibed seeds has been found effective for breaking dormancy in many species. e.g. brinjal, Malus sylvestris, etc. 5. Washing: Washing of dormant seeds by water has been reported to break dormancy, probably due to leaching of dormancy factor from the seeds. e.g. ragi, groundnut, etc.
  • 20. 6. Dry storage: The embryo of seeds of certain species is not fully developed even at harvestable maturity stage and requires some more time after harvest (after ripening period) for full development of embryo. Dry storage of such seeds for few days will break dormancy and promote germination. 7. Chemical treatments: Various chemicals and growth hormones, viz. Gibberellins, cytokinins, ethylene, KNO3, thiourea, HNO3, cyanide, etc. have been found to break seed dormancy in different species. The concentration of the chemicals and mode of treatment varies in seeds. Polymorphism of dormancy: In sunflower, the seeds formed in the peripheral portion of the head/capitulum may not be dormant while those formed in the central portion are dormant. This is known as polymorphism in seed dormancy.
  • 21. Classification of fruit crops Classification is the system of grouping or placing of individuals according to nomenclature. It is very useful to the pomologist. It helps to:  To identify and naming the crop.  To study the close relationship.  To know their hybrids and crossing behaviour.  To know their compatibility & inter grafting ability.  To know their adoptability to soil & climate. POMOLOGY:  Pomology is a branch of horticulture which deals with study of various aspects of fruits loke, rising of saplings, growing them properly and providing various intercultural operations.  The term pomology is a combination of two Latin words „Pome‟ means ‗Fruits‟ and ‗Logos„ means ‗study‟.  “Poma” in Greek also meaning fruits later subsequently “Pome” in Latin word means fruits, logosstudy.  Basic Pomology: Study of basic aspects of fruit production like training, water management, use of PGR„s.  Commercial Pomology: It is concerned with commercial production of fruits.  Systematic Pomology: It may be concerned with classification and nomenclature like kingdom, order, class, genus and species. A. Classification of fruits based on climate adaptability. In this classification, the fruits trees are categorized into three recognized groups. i. Temperate fruits:  Temperate fruit plants are exacting in their climate requirement.  They are grown only in place where winter is distinctly cold, require as exposure of specific chilling temperature for certain period without which they do not flower.  These fruit plants are generally deciduous and stand frost. Eg. Apple, almond, peach, pear, plum, strawberry, apricot, persimmon, cherymoya, walnut, peanut, hassle nut, cherry, pistachios and kiwifruits etc. ii. Tropical fruits:  Tropical fruit plants are generally evergreen and are extremely sensitive to cold.
  • 22.  They do well under lesser fluctuations of diurnal temperature, light and dark periods they require a moist warm climate but are capable of withstanding dry weather in some cases Eg; mango, banana, papaya, sapota, etc., iii. Sub-tropical fruits:  The fruit crops grown under a climatic condition between temperate and the tropical are known as subtropical fruit crops. They may be either deciduous or evergreen and are usually able to withstand a low temperature but not the frost.  They are also quite adoptive to fluctuations of light and dark period during day and night.  Some subtropical fruit plants require chilling for flower bud differentiation.  Example; grape, citrus, durian, jackfruit, etc., B. Classification based on bearing habit: On the basis of bearing habit, fruit trees are classified in to six categories to facilitate cultural operation like pruning, skiffing, heading back etc. 1. Fruit buds bore terminally and giving rise to inflorescence without leaves e.g. Mango, Cherry, etc. 2. Fruit buds borne terminally and unfolding to produce leafy shoots which terminate in flower clusters. e.g. Apple 3. Fruit buds borne terminally and unfolding to produce leafy shoots with flower or flower clusters e.g Guava 4. Fruit bud borne laterally containing flower parts only and giving rise to inflorescence without leaves or leaves present, they are reduced in size., e.g. Citrus 5. Fruit bud borne laterally and unfolding to produce leafy shoots terminally in flower clusters this type of flowering is noticed in grapes and cashewnut. 6. Fruit buds borne laterally and unfolding to produce leafy shoots with flower clusters in leafy axils. eg.Fig. C. Fruit morphology: 1) Simple fruit - Berry: Banana, Papaya, Grape, Sapota, and Avocado 2) Modified berry i. Balusta : Pomegranate ii. Amphisarca : Woodapple, Bael
  • 23. iii. Pepo : Water melon iv. Pome : Apple, Pear, Laquat v. Drupe (Stone) : Mango, Pear, Plum vi. Hesperidium : Citrus vii. Nut : Cashew, Litchi, Walnut, Rambutan viii. Capsule : Anola, Carambola 3) Aggregate fruits : Etario of berries –Custard apple, Raspberry 4) Multiple fruit : Syconus- Fig : Sorosis- Jackfruit, Pineapple, Mulberry D. Based on rate of respiration: Climacteric Fruits Non-climacteric Fruits Mango, Banana, Sapota, Guava, Papaya, Apple, Fig, Peach, Pear, Plum, Annona, Tomato Citrus, Grape, Pomegranate Pineapple Litchi, Ber, Jamun, Cashew, Cucumber, Cherry, Strawberry (Climacteric fruits produce much larger amount of ethylene than non climacteric fruits) E. Based on photoperiodic responses Long day Short day Day neutral plant Passionfruit, Banana, Apple Strawberry, Pineapple, Coffee Papaya, Guava F. Based on relative salt tolerance Highly tolerant Medium tolerant Highly sensitive Datepalm, Ber, Amla, Guava, Coconut, Khirni Pomegranate, Cashew, Fig, Jamun, Phalsa Mango, Apple, Citrus, Pear, Straw berry G. Based on relative acid Tolerance Highly tolerant Medium tolerant Highly sensitive Stawberry, Raspberry, Fig, Bael, Plum Pineapple, Avocado, Litchi H. Based on longevity: a) Very Long longevity - >100yrs- Datepalm, Coconut, Arecanut b) Long longevity - 50-100yrs - Mango, Tamarind c) Medium - 10-50yrs - Litchi, Guava,Pomogranate d) Short - Pineapple, Banana I. Based on consumers preference or weight of fruits
  • 24. a) Very light - 50-100gm Grape,Ber,Banana b) Light - 100-150gm Sapota,Pomegranate c) Light medium - 150-300gm Mango d) Medium - 300-350gm Avocado e) Medium to heavy - 800-1000gm Mango f) Heavy - 1-5kg Bread fruit, Pineapple g) Very heavy - >5kg Jack Fruit Area, production, productivity and export potential