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Agriculture and Natural Resources

                                                                                                                                   FSA6024




                          Plant Propagation

                         for Home Gardeners

James A. Robbins                            Many home gardeners wish to                             sterile and typically holds too much
Professor and                          reproduce special plants in their land­                      water. Manufacturers often sell a soil­
                                       scapes or start their own vegetable                          less media designed specifically for
Horticulture Specialist ­              and bedding plants. Plant propagation                        propagation. The media should be
Ornamentals                            is the controlled reproduction of                            moist before sowing the seed. For very
                                       plants using a variety of methods.                           small seeds, it may be easier to
                                       These methods can be broadly                                 sprinkle the seeds on the media
                                       grouped into sexual and asexual                              surface and then use a fine spray of
                                       (vegetative) methods. Sexual propa­                          water to settle them into the media.
                                       gation is focused primarily on                               For larger seeds, you can sow the
                                       propagating plants by seed. Seed                             seeds and then cover them with an
                                       propagation offers some advantages in                        appropriate depth of moistened media.
                                       that the technique is often economical                       The general rule of thumb for covering
                                       and efficient and lends itself to long-                      seeds is to cover the seed with two
                                       term storage. The primary disadvan­                          times the diameter/width of the
                                       tage of seed propagation is that many                        seed. Again, use water to settle the
                                       seeds do not come “true to type,” so                         media in after topdressing the seed
                                       the offspring are not exact copies of                        with media.
                                       the parent. Asexual propagation tech­
                                       niques include cuttings (e.g., root,
                                       stem, leaf), budding or grafting, layer­
                                       ing and division. Asexual propagation
                                       techniques, while often more labor
                                       intensive than seed propagation, do
                                       allow the gardener to get an exact
                                       copy of the parent plant.

                                       Seed Propagation
                                           Seed propagation is commonly
                                       used by a home gardener when
                                       starting plants for the vegetable                             Figure 1. Germinating seed.
                                       garden or for some bedding plants.
                                       Most gardeners are well aware of how
                                       to prepare a seedbed in a vegetable                               Keeping the media moist (i.e., not
                                       garden and then follow the directions                        wet and not dry) during germination
                                       on the seed packet to sow the seeds.                         of the seed is critical. Once the media
                                                                                                    is at the correct moisture level, simple
                                            Starting seeds indoors for the                          techniques can be used to maintain
                                       vegetable garden or for bedding plants                       that level for the entire germination
                                       is fairly simple. The two most critical                      period. One simple method involves
                                       factors are using a high-quality seed                        placing a glass plate or sheet of plas­
Arkansas Is
                           germination mix and then keeping the                         tic or Saran™ above the lip of the
                                       seeds moist but not overly wet. Media                        seed tray or container. If the plate or
Our Campus
                            for seed germination should ideally be                       sheet is elevated just slightly above
                                       sterile, most likely consist of a                            the media surface, it will need to be
                                       significant percentage of peat moss                          raised or removed once the seeds
Visit our web site at:                 and never contain garden soil. Garden                        begin to germinate and grow. The
http://www.uaex.edu                    soil is a poor choice because it is not                      other common option is to place a

                         University of Arkansas, United States Department of Agriculture, and County Governments Cooperating
clear plastic bag over the container, again with the        surface. Seeds can also be scarified using hot water
goal of simply trapping existing moisture contained in      (170 to 212 degrees F). With this method, simply drop
the media. If the bag is not rigid, it is common to use     the seeds into four to five times their volume of hot
an appropriate length of bamboo stake or chopstick to       water, remove water from the heat source and let the
keep the bag from collapsing in on the fragile              seeds soak for 12 to 24 hours unless the literature
seedlings. Some seedlings are very sensitive to salts       states otherwise.
in water; therefore, for small seed germination
projects, you may wish to use deionized bottled water
until seedlings are up and actively growing. If seed        Vegetative Propagation
has been sown with a high density, you will most
likely have to transplant the fragile seedlings to a            Vegetative (asexual) propagation techniques
different container (e.g., cell pack) to allow the          include cuttings (e.g., root, stem, leaf), budding or
seedlings to reach a size that can be transplanted into     grafting, layering and division. Propagation tech­
the garden or landscape.                                    niques such as cuttings, layering and division can be
                                                            easily performed by home gardeners; however,
                                                            budding and grafting typically require more skill.

                                                            Cuttings

                                                                 An understanding
                                                            of terminology is
                                                            required before going
                                                            further. A number of
                                                            terms are used when
                                                            discussing cutting
                                                            propagation. Simple
                                                            terms are used to
                                                            describe where the
                                                            cutting originated
Figure 2. Seed germination trays.                           from (e.g., root, stem, Figure 3. Typical three-node
                                                            leaf), or with stem       semi-hardwood cutting with
     Especially with woody plants, you will need to         cuttings, the relative basal leaves removed.
become familiar with two terms: stratification and          age of the wood/tissue
scarification. In a simple sense, stratification involves   (e.g., softwood, semi-hardwood, hardwood). With stem
different methods imposed by man to cause seeds to          cuttings, we may also refer to the number of nodes.
germinate after simulating environmental conditions         Nodes are simply the locations on the stem where the
that seeds would normally experience in nature. For         leaves originate from (i.e., more precisely, where the
example, think about a red oak acorn that falls to the      buds are located). It is also common to refer to the
ground in the fall. In nature it is exposed to winter       overall position on the stem that the cutting was
temperatures and then begins germinating the                taken from (e.g., terminal, sub-terminal). For exam­
following spring. The internal signals keep the acorn       ple, it is common for a gardening book to instruct you
from germinating in the fall before harsh weather           to collect “terminal cuttings with three to four nodes.”
sets in. Knowing this, if you harvest red oak acorns in     With most cuttings, you also need to pay attention to
the fall and then want to germinate the seed indoors,       the original “up and down” ends of your cuttings
you need to replicate what would have gone on in            (polarity). If you think you will have trouble keeping
nature. Therefore, the red oak acorns are placed in a       track once you have removed a cutting from a stem,
container with moistened media and then placed in a         you can simply use a lateral or cross-cut to reference
refrigerator (32 to 45 degrees F) for three months.         the top/“up” and an angle cut to reference the
These “stratified” seeds can then be removed from the       bottom/“down” of the cutting.
refrigerator and sown, and germination should begin
soon if the temperatures are suitable. Reference                The media you use to root cuttings can have a
books will clearly mention what kind of stratification      significant impact on your rooting results, not only
regime is required for each species.                        the number of new roots but the growth of these
                                                            roots. An ideal rooting media needs to be sterile, well
    Scarification is any physical process that softens      drained and provide adequate oxygen. Individual
or damages the seed coat making it more permeable           components that are most commonly used include
to water and air. Again, propagation books clearly          coarse perlite, coarse vermiculite, peat moss and
indicate which species require scarification to             sand/pumice. These individual components can be
improve germination. Plants in the Legume family            combined in a variety of ways and percentages.
are one group often mentioned. For small seeds, you         Examples include 50% peat moss: 50% perlite
can simply place the seeds between two layers of            (volume basis); 50% perlite: 50% vermiculite; 100%
medium sand paper and then rub the sheets back and          perlite; 100% sand/pumice; and 100% peat moss.
forth. For larger seeds (e.g., Kentucky coffee tree),       Rooting cubes (e.g., Oasis Rootcubes®) made out of
you can use a metal or nail file to abrade the seed         foam are also used.
have formed roots, you can start this transition by
                                                             periodically venting your cover with increasing
                                                             frequency to reduce the relative humidity slowly
                                                             over time.




Figure 4. Effect of media on rooting performance.
Left: vermiculite, peat moss, perlite, peat:perlite

     The rooting environment has a major influence
on the rooting results. Remember that with cuttings
we have separated a portion of the plant from the
parent plant, thus removing its source of water and
nutrients until a new root system has reestablished.         Figure 5. Simple propagation option using plastic bag.
Environmental conditions that add stress or hasten
water loss diminish the rooting success. Therefore,
conditions such as high light, high temperature or
breezy conditions often have a negative impact on the
rooting results. Even with cuttings that have leaves,
it is hard for people to imagine that making “food” is
not a priority, but forming roots is. Cuttings will rely
on stored “food” until new roots are regenerated. With
this in mind, ideal temperature conditions for most
cuttings are to have the air temperature 10 to 15
degrees cooler than the rooting media. Ideal tempera­
tures for many types of cuttings would be an ambient
temperature of 55 to 65 and a media temperature of
65 to 75 degrees F. In most home situations, having a
dual temperature is difficult; however, garden               Figure 6. High dome for rooting cuttings.
suppliers do sell waterproof heating mats designed
specifically to place propagation trays on to elevate
the media temperature. As for light levels, it is criti­
cal to keep cuttings out of direct sunlight until they
have rooted. Appropriate light levels can be found in
north-facing windows or by using grow lights in dark
rooms or basements.

     The critical factor with cuttings that contain
leaves is to maintain the highest level of relative
humidity possible around leaves until roots form.
Commercial propagators use intermittent mist sys­
tems or fog systems to achieve this goal. While these        Figure 7. Simple propagation tray.
options are possible for a homeowner, there are easier
solutions. No matter what option you chose, the goal
is the same: you are trying to trap moisture already
present in your rooting media. For very few cuttings,
using an inverted Ziploc™ bag or one-half of a clear
plastic soda bottle works very well over a single pot.
If you use the threaded end of a plastic bottle, you
can easily remove the bottle cap to vent moisture if it
builds up too high in the bottle. For large numbers of
cuttings, it is easier to use a seed tray or flat. To trap
moisture when using a tray, use a clear tray lid or
pieces of bent strap metal (Fig. 7) to keep a plastic
bag from touching the cuttings. With most leafy
cuttings, we need to transition or “harden off” cut­
tings before planting them outside. Once cuttings            Figure 8. Plastic bottles (top or bottom) for rooting cuttings.
Houseplants                                                (lamina) and the petiole that attaches it to the
                                                           original stem. This method is commonly used for
    Many different kinds of common houseplants can         African violet, peperomia and some begonias. Roots
be easily propagated by stem, cane or leaf cuttings        and a new plantlet typically originate at the base of
and by division or layering.                               the petiole.

Stem Cuttings                                                  Leaf-Blade Cuttings – Leaf-blade cuttings consist
                                                           of the leaf blade only and are commonly used to
    There are a number of common houseplants that          propagate many succulent plants such as jade plant
can be easily propagated by stem cuttings. These           and sedums. In this case the leaf is most commonly
include coleus, begonia, Christmas cactus, fuchsia,        inserted vertically into the media with a majority of
geranium, poinsettia, rubber plant, shrimp plant and       the leaf visible above the media surface. Roots and
wandering jew. Take cuttings from shoots that are          new plantlets typically originate from the base of the
vigorous, pest free and, ideally, not flowering.           leaf. One exception would be kalanchoe. In this case
Cuttings will typically be 4 to 6 inches long depend­      you lay the leaf horizontally on the media surface
ing on the plant. Cut the stem just below a node with      and plantlets will develop at many of the serrations
a sharp knife or razor blade. Remove leaves from the       or notches in the margin of the leaf.
lower half of the cutting and dip the basal 1 to 1.5
inches in a rooting hormone. Rooting hormone is                Leaf-Vein Cuttings – The classic example of this
probably not necessary for easy-to-root plants such as     cutting method is the begonia. You simply lay the leaf
coleus and fuchsia, but it will not hinder the rooting     on the media surface and cut several slices perpen­
process either. Carefully stick the dipped end of the      dicular to the major leaf veins. Some people also flip
stem cutting into the rooting media. Some people like      the leaf over and lightly dust those cuts before laying
to pre-drill the hole using the eraser end of a pencil.    the leaf back on the media surface. Roots and
The rooting media would then be firmed around the          plantlets will develop at the cuts in the veins.
cutting after inserting into this preformed hole.
                                                               Leaf-Section Cuttings – The most common
    A modification of this works well for foliage          example of this technique is used to propagate
plants that have a larger stem such as dumb cane,          mother-in-law’s tongue (Sansevieria). In this case you
corn plant and Chinese evergreen. In this case you         cut the long “tongue-like” leaf into 2-inch segments
will remove a leafless section of the stem, now            and insert the bottom ½ inch into the rooting media.
referred to as a cane, and lay this cane horizontally      Remember to keep the original orientation of the
on the media surface. If the cane is thick, you will       leaf intact (polarity). Some of the baby plantlets that
push the cane slightly into the media surface so you       will form at the bottom of the leaf piece from a
get good media contact.                                    variegated selection may develop as green only (i.e.,
                                                           no variegation).
Leaf Cuttings

    Leaf cuttings are used to propagate many house­
plants including begonia, African violet, Episcia, glox­
inia, jade plant, kalanchoe, mother-in-law’s tongue
and peperomia. There are several variations of leaf
cutting types depending on what parts of the leaf are
used and whether a leaf petiole is attached or not.
    Leaf-Petiole Cuttings – Just as it sounds, a leaf-
petiole cutting includes both the wide part of the leaf




                                                           Figure 10. Mother-in-law’s tongue leaf-section cuttings.


                                                               Preformed Plantlets – A small number of house­
                                                           plants produce preformed plantlets along aerial
                                                           shoots. Examples include spider plant, wax plant
                                                           (Hoya) and strawberry begonia (Saxifraga
                                                           stolonifera). Propagation is simple involving the
                                                           removal of these plantlets (with or without aerial
                                                           rootlets) and then inserting the base of the plantlets
                                                           in a high-quality potting soil and treating the
Figure 9. African violet leaf-petiole cuttings.            plantlet as if it were a stem cutting.
Outdoor Plants
    Many of your favorite outdoor shrubs, vines and
ground covers can be easily propagated by cuttings.

Softwood Cuttings
    Softwood cuttings are prepared from the soft,
succulent, new flush of growth. Many ornamental
plants can be propagated by softwood cuttings includ­
ing lilacs, forsythia, magnolia, weigela, oleander,
pyracantha, spirea, apple, peach, pear and plum.
While softwood cuttings tend to root easily and faster,
they may require more care since the soft tissue is
more susceptible to drying. In general, with softwood     Figure 12. Hardwood cutting.
cuttings, we are talking about cuttings that are
typically 3 to 4 inches long, with two to three nodes
and the basal leaves removed before inserting the         Root Cuttings
cutting into the media.                                        Root cuttings are used very rarely and only on a
                                                          few species that typically do not root well from stem
Semi-Hardwood Cuttings                                    cuttings. These include Aesculus, Aralia,
    Realistically, terms such as softwood and semi-       Koelreuteria, sumac (Rhus) and sassafras. The most
hardwood are not absolutes. Once the very soft flush      challenging aspect of root cuttings is uncovering from
of new growth starts to stiffen up and form “wood,”       soil the roots of your desired plant and not some
we have likely passed into the semi-hardwood stage.       other plant. While root cuttings may be made at any
A majority of our common ornamental shrubs are            time of the year, it is probably more common to do
propagated from semi-hardwood cuttings. Plants such       this in the dormant season. Root pieces can be held in
as viburnum, camellia, azalea, holly, crape myrtle,       moist media in a cooler (>38 degrees), placed nearly
hydrangea and euonymus use this type of cutting.          horizontal in rooting media like you would treat other
In Arkansas this peak time is often between May 15        cuttings or direct planted in well-prepared garden
and June 15.                                              soil with the top of the root-piece just barely
                                                          emerging from the soil.

                                                          Grafting/Budding
                                                              Realistically, grafting or budding is not an easy
                                                          technique for home gardeners to master. Most people
                                                          would be amazed at how many ornamental plants in
                                                          their landscape originated from budded or grafted
                                                          plants. Many shade trees, all Japanese maples, many
                                                          weeping plants and all hybrid tea roses use these
                                                          techniques. Budding or grafting is also the mainstay
                                                          of the tree fruit industry. Budding and grafting are
                                                          essentially the same technique; however, with
                                                          budding you are simply attaching the bud from your
                                                          desired plant to the rootstock/understock, and with
Figure 11. Dipping prepared holly cutting in rooting      grafting you are using a piece of stem that contains
hormone.                                                  many buds on it. The obvious advantage of bud­
                                                          ding is you can potentially get many more plants
Hardwood Cuttings                                         from your desired scion/stockplant. The major
    Hardwood cuttings are made from woody,
dormant wood on deciduous plants (e.g., grape,
poplar, rose, willow, peach, wisteria, elderberry,
blackberry, forsythia) or some needle evergreens (e.g.,
junipers, arborvitae). Hardwood cuttings are easy to
prepare and less perishable. For deciduous plants the
cuttings are leafless and tend to be longer than soft­
wood cuttings. Hardwood cuttings are typically 6 to
12 inches long. As a general rule for the needle ever­
greens, rooting percentages tend to be increased
when cuttings are harvested in mid- to late winter.
As a general rule, higher rooting percentages occur
on junipers when a small “heal” of wood is included
at the base of the cutting.                               Figure 13. Chip budding.
disadvantage of budding is that you are typically                                for plants such as wisteria, brambles, forsythia,
working with a very small bud versus a larger piece                              weigela and spirea. Ground layering simply involves
of stem that is easier to handle. A good propagation                             bending an intact shoot over and burying the middle
text can go into the specific details related to budding                         portion (i.e., shoot tip comes back up above ground
and grafting.                                                                    and is exposed) under the existing soil. You may wish
                                                                                 to wound the lower portion of the bent stem. With
Layering                                                                         easy-to-root plants, roots will often form along the
    Layering is a method of rooting a new plant while                            buried stem. Once roots have formed, you simply cut
the stem is still attached to the parent plant. It is                            the stem below the rooted zone, excising the plant
simple to perform and can be done in the home                                    from the mother plant.
without special equipment or
structures.                                                                      Division
     Air Layering – In the                                                            Many home gardeners did not realize that they
case of air layering, we are                                                     were actually plant propagators when they divided
forming roots on stems well                                                      common perennials such as cannas, hosta, daylilies,
above the soil. This method                                                      iris, peonies, bleeding heart, many ornamental
can be used successfully on a                                                    grasses, ferns and lily-of-the-valley. As a general rule,
number of common tropical                                                        the best time to divide plants is during the dormant
houseplants including croton,                                                    season.
dumb cane (Dieffenbachia),
dracaena, rubber plant and
schefflera.                                                                      Resources
     The process is quite                                                        Bubel, Nancy. 1988. The New Seed Starter’s
simple. First, pick a point in                                                       Handbook. ISBN-10:0878577521.
                                   Figure 14. Air layering
the stem at least 10 to 12         Mahonia.                                      Dirr, M.A., and C.W. Heuser, J. 2006. The Reference
inches from the tip. Remove                                                          Manual of Woody Plant Propagation: From Seed
all leaves several inches on either side of the region                               to Tissue Culture: A Practical Working Guide to
where you will make your air layer. The area will                                    the Propagation of Over 1,100 Species. Varsity
typically be about 6 inches in length. Two types of                                  Press. ISBN-10:0942375092. Basically an A to Z
stem cuts are typically used. One involves making an                                 recipe book for propagating woody ornamental
upward-angled cut into the stem that stops about                                     plants.
halfway through the stem. The other method involves                              Druse, Ken. 2000. Making More Plants: The Science,
removing a ring of the outer bark. The ring of bark                                  Art and Joy of Propagation. $32. ISBN-10:
tissue will be 1 to 1.5 inches long. Apply a rooting                                 051770787X.
hormone to the cut surfaces. If an angled or vertical                            Gowdy, Mary Ann. 2002. Home Propagation of House­
cut has been made into the stem, you will want to                                    plants. University of Missouri - Columbia. G6560.
prevent that from closing by inserting a small piece                                 http://extension.missouri.edu/explorepdf/
of toothpick in the wounded area. Next, take a hand­                                 agguides/hort/g06560.pdf.
ful of moist sphagnum moss and pack the moss in                                  Hartmann, H.T., D.E. Kester, F.T. Davies, Jr., and
and around the cut surfaces. To retain the moisture                                  R.L. Geneve. 2001. Plant Propagation Principles
around the wounded area for weeks/months, wrap a                                     and Practices. Prentice-Hall. 7th edition. ISBN­
layer of plastic sheet around the entire work area                                   10:0136792359. This is considered the bible of
and seal both ends with electrical tape. Many people                                 plant propagation. It is used in every college and
also cover the plastic with aluminum foil to prevent                                 university. Expensive, but thorough.
any light from penetrating the rooting zone and to
                                                                                 Heffernan, Maureen. 1997. Burpee Seed Starter. John
reflect any heat from sunlight. You can simply keep
                                                                                     Wiley & Sons. ISBN-10:0028610024.
your foliage plant in the same place (i.e., indoors or
outdoors) where it was originally growing. Once roots
do form, simply make a single cross-cut in the stem
                                                                                 Supplies
below the rooting zone. You should end up with roots                             •	 Your local garden supply store can often provide
on the bottom and leaves at the top: a new plant.                                   trays, media and rooting hormone.
   Ground Layering – Ground layering can be                                      •	 A.M. Leonard, Piqua, OH, http://amleo.com/
used to easily propagate a number of ornamental                                  •	 Hummert International, Earth City, MO,
shrubs and vines. This simple technique works well                                  http://www.hummert.com/
                   No endorsement is implied or discrimination intended for firms or references included or excluded from this publication.

                                   Printed by University of Arkansas Cooperative Extension Service Printing Services.


DR. JAMES A. ROBBINS is professor and horticulture specialist -                  Issued in furtherance of Cooperative Extension work, Acts of May 8
ornamentals, University of Arkansas Division of Agriculture,                     and June 30, 1914, in cooperation with the U.S. Department of
Cooperative Extension Service, Little Rock.                                      Agriculture, Director, Cooperative Extension Service, University of
                                                                                 Arkansas. The Arkansas Cooperative Extension Service offers its
                                                                                 programs to all eligible persons regardless of race, color, national
                                                                                 origin, religion, gender, age, disability, marital or veteran status, or
                                                                                 any other legally protected status, and is an Affirmative
                                                     FSA6024-PD-4-08RV           Action/Equal Opportunity Employer.

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Plant Propagation for Home Gardeners

  • 1. Agriculture and Natural Resources FSA6024 Plant Propagation for Home Gardeners James A. Robbins Many home gardeners wish to sterile and typically holds too much Professor and reproduce special plants in their land­ water. Manufacturers often sell a soil­ scapes or start their own vegetable less media designed specifically for Horticulture Specialist ­ and bedding plants. Plant propagation propagation. The media should be Ornamentals is the controlled reproduction of moist before sowing the seed. For very plants using a variety of methods. small seeds, it may be easier to These methods can be broadly sprinkle the seeds on the media grouped into sexual and asexual surface and then use a fine spray of (vegetative) methods. Sexual propa­ water to settle them into the media. gation is focused primarily on For larger seeds, you can sow the propagating plants by seed. Seed seeds and then cover them with an propagation offers some advantages in appropriate depth of moistened media. that the technique is often economical The general rule of thumb for covering and efficient and lends itself to long- seeds is to cover the seed with two term storage. The primary disadvan­ times the diameter/width of the tage of seed propagation is that many seed. Again, use water to settle the seeds do not come “true to type,” so media in after topdressing the seed the offspring are not exact copies of with media. the parent. Asexual propagation tech­ niques include cuttings (e.g., root, stem, leaf), budding or grafting, layer­ ing and division. Asexual propagation techniques, while often more labor intensive than seed propagation, do allow the gardener to get an exact copy of the parent plant. Seed Propagation Seed propagation is commonly used by a home gardener when starting plants for the vegetable Figure 1. Germinating seed. garden or for some bedding plants. Most gardeners are well aware of how to prepare a seedbed in a vegetable Keeping the media moist (i.e., not garden and then follow the directions wet and not dry) during germination on the seed packet to sow the seeds. of the seed is critical. Once the media is at the correct moisture level, simple Starting seeds indoors for the techniques can be used to maintain vegetable garden or for bedding plants that level for the entire germination is fairly simple. The two most critical period. One simple method involves factors are using a high-quality seed placing a glass plate or sheet of plas­ Arkansas Is germination mix and then keeping the tic or Saran™ above the lip of the seeds moist but not overly wet. Media seed tray or container. If the plate or Our Campus for seed germination should ideally be sheet is elevated just slightly above sterile, most likely consist of a the media surface, it will need to be significant percentage of peat moss raised or removed once the seeds Visit our web site at: and never contain garden soil. Garden begin to germinate and grow. The http://www.uaex.edu soil is a poor choice because it is not other common option is to place a University of Arkansas, United States Department of Agriculture, and County Governments Cooperating
  • 2. clear plastic bag over the container, again with the surface. Seeds can also be scarified using hot water goal of simply trapping existing moisture contained in (170 to 212 degrees F). With this method, simply drop the media. If the bag is not rigid, it is common to use the seeds into four to five times their volume of hot an appropriate length of bamboo stake or chopstick to water, remove water from the heat source and let the keep the bag from collapsing in on the fragile seeds soak for 12 to 24 hours unless the literature seedlings. Some seedlings are very sensitive to salts states otherwise. in water; therefore, for small seed germination projects, you may wish to use deionized bottled water until seedlings are up and actively growing. If seed Vegetative Propagation has been sown with a high density, you will most likely have to transplant the fragile seedlings to a Vegetative (asexual) propagation techniques different container (e.g., cell pack) to allow the include cuttings (e.g., root, stem, leaf), budding or seedlings to reach a size that can be transplanted into grafting, layering and division. Propagation tech­ the garden or landscape. niques such as cuttings, layering and division can be easily performed by home gardeners; however, budding and grafting typically require more skill. Cuttings An understanding of terminology is required before going further. A number of terms are used when discussing cutting propagation. Simple terms are used to describe where the cutting originated Figure 2. Seed germination trays. from (e.g., root, stem, Figure 3. Typical three-node leaf), or with stem semi-hardwood cutting with Especially with woody plants, you will need to cuttings, the relative basal leaves removed. become familiar with two terms: stratification and age of the wood/tissue scarification. In a simple sense, stratification involves (e.g., softwood, semi-hardwood, hardwood). With stem different methods imposed by man to cause seeds to cuttings, we may also refer to the number of nodes. germinate after simulating environmental conditions Nodes are simply the locations on the stem where the that seeds would normally experience in nature. For leaves originate from (i.e., more precisely, where the example, think about a red oak acorn that falls to the buds are located). It is also common to refer to the ground in the fall. In nature it is exposed to winter overall position on the stem that the cutting was temperatures and then begins germinating the taken from (e.g., terminal, sub-terminal). For exam­ following spring. The internal signals keep the acorn ple, it is common for a gardening book to instruct you from germinating in the fall before harsh weather to collect “terminal cuttings with three to four nodes.” sets in. Knowing this, if you harvest red oak acorns in With most cuttings, you also need to pay attention to the fall and then want to germinate the seed indoors, the original “up and down” ends of your cuttings you need to replicate what would have gone on in (polarity). If you think you will have trouble keeping nature. Therefore, the red oak acorns are placed in a track once you have removed a cutting from a stem, container with moistened media and then placed in a you can simply use a lateral or cross-cut to reference refrigerator (32 to 45 degrees F) for three months. the top/“up” and an angle cut to reference the These “stratified” seeds can then be removed from the bottom/“down” of the cutting. refrigerator and sown, and germination should begin soon if the temperatures are suitable. Reference The media you use to root cuttings can have a books will clearly mention what kind of stratification significant impact on your rooting results, not only regime is required for each species. the number of new roots but the growth of these roots. An ideal rooting media needs to be sterile, well Scarification is any physical process that softens drained and provide adequate oxygen. Individual or damages the seed coat making it more permeable components that are most commonly used include to water and air. Again, propagation books clearly coarse perlite, coarse vermiculite, peat moss and indicate which species require scarification to sand/pumice. These individual components can be improve germination. Plants in the Legume family combined in a variety of ways and percentages. are one group often mentioned. For small seeds, you Examples include 50% peat moss: 50% perlite can simply place the seeds between two layers of (volume basis); 50% perlite: 50% vermiculite; 100% medium sand paper and then rub the sheets back and perlite; 100% sand/pumice; and 100% peat moss. forth. For larger seeds (e.g., Kentucky coffee tree), Rooting cubes (e.g., Oasis Rootcubes®) made out of you can use a metal or nail file to abrade the seed foam are also used.
  • 3. have formed roots, you can start this transition by periodically venting your cover with increasing frequency to reduce the relative humidity slowly over time. Figure 4. Effect of media on rooting performance. Left: vermiculite, peat moss, perlite, peat:perlite The rooting environment has a major influence on the rooting results. Remember that with cuttings we have separated a portion of the plant from the parent plant, thus removing its source of water and nutrients until a new root system has reestablished. Figure 5. Simple propagation option using plastic bag. Environmental conditions that add stress or hasten water loss diminish the rooting success. Therefore, conditions such as high light, high temperature or breezy conditions often have a negative impact on the rooting results. Even with cuttings that have leaves, it is hard for people to imagine that making “food” is not a priority, but forming roots is. Cuttings will rely on stored “food” until new roots are regenerated. With this in mind, ideal temperature conditions for most cuttings are to have the air temperature 10 to 15 degrees cooler than the rooting media. Ideal tempera­ tures for many types of cuttings would be an ambient temperature of 55 to 65 and a media temperature of 65 to 75 degrees F. In most home situations, having a dual temperature is difficult; however, garden Figure 6. High dome for rooting cuttings. suppliers do sell waterproof heating mats designed specifically to place propagation trays on to elevate the media temperature. As for light levels, it is criti­ cal to keep cuttings out of direct sunlight until they have rooted. Appropriate light levels can be found in north-facing windows or by using grow lights in dark rooms or basements. The critical factor with cuttings that contain leaves is to maintain the highest level of relative humidity possible around leaves until roots form. Commercial propagators use intermittent mist sys­ tems or fog systems to achieve this goal. While these Figure 7. Simple propagation tray. options are possible for a homeowner, there are easier solutions. No matter what option you chose, the goal is the same: you are trying to trap moisture already present in your rooting media. For very few cuttings, using an inverted Ziploc™ bag or one-half of a clear plastic soda bottle works very well over a single pot. If you use the threaded end of a plastic bottle, you can easily remove the bottle cap to vent moisture if it builds up too high in the bottle. For large numbers of cuttings, it is easier to use a seed tray or flat. To trap moisture when using a tray, use a clear tray lid or pieces of bent strap metal (Fig. 7) to keep a plastic bag from touching the cuttings. With most leafy cuttings, we need to transition or “harden off” cut­ tings before planting them outside. Once cuttings Figure 8. Plastic bottles (top or bottom) for rooting cuttings.
  • 4. Houseplants (lamina) and the petiole that attaches it to the original stem. This method is commonly used for Many different kinds of common houseplants can African violet, peperomia and some begonias. Roots be easily propagated by stem, cane or leaf cuttings and a new plantlet typically originate at the base of and by division or layering. the petiole. Stem Cuttings Leaf-Blade Cuttings – Leaf-blade cuttings consist of the leaf blade only and are commonly used to There are a number of common houseplants that propagate many succulent plants such as jade plant can be easily propagated by stem cuttings. These and sedums. In this case the leaf is most commonly include coleus, begonia, Christmas cactus, fuchsia, inserted vertically into the media with a majority of geranium, poinsettia, rubber plant, shrimp plant and the leaf visible above the media surface. Roots and wandering jew. Take cuttings from shoots that are new plantlets typically originate from the base of the vigorous, pest free and, ideally, not flowering. leaf. One exception would be kalanchoe. In this case Cuttings will typically be 4 to 6 inches long depend­ you lay the leaf horizontally on the media surface ing on the plant. Cut the stem just below a node with and plantlets will develop at many of the serrations a sharp knife or razor blade. Remove leaves from the or notches in the margin of the leaf. lower half of the cutting and dip the basal 1 to 1.5 inches in a rooting hormone. Rooting hormone is Leaf-Vein Cuttings – The classic example of this probably not necessary for easy-to-root plants such as cutting method is the begonia. You simply lay the leaf coleus and fuchsia, but it will not hinder the rooting on the media surface and cut several slices perpen­ process either. Carefully stick the dipped end of the dicular to the major leaf veins. Some people also flip stem cutting into the rooting media. Some people like the leaf over and lightly dust those cuts before laying to pre-drill the hole using the eraser end of a pencil. the leaf back on the media surface. Roots and The rooting media would then be firmed around the plantlets will develop at the cuts in the veins. cutting after inserting into this preformed hole. Leaf-Section Cuttings – The most common A modification of this works well for foliage example of this technique is used to propagate plants that have a larger stem such as dumb cane, mother-in-law’s tongue (Sansevieria). In this case you corn plant and Chinese evergreen. In this case you cut the long “tongue-like” leaf into 2-inch segments will remove a leafless section of the stem, now and insert the bottom ½ inch into the rooting media. referred to as a cane, and lay this cane horizontally Remember to keep the original orientation of the on the media surface. If the cane is thick, you will leaf intact (polarity). Some of the baby plantlets that push the cane slightly into the media surface so you will form at the bottom of the leaf piece from a get good media contact. variegated selection may develop as green only (i.e., no variegation). Leaf Cuttings Leaf cuttings are used to propagate many house­ plants including begonia, African violet, Episcia, glox­ inia, jade plant, kalanchoe, mother-in-law’s tongue and peperomia. There are several variations of leaf cutting types depending on what parts of the leaf are used and whether a leaf petiole is attached or not. Leaf-Petiole Cuttings – Just as it sounds, a leaf- petiole cutting includes both the wide part of the leaf Figure 10. Mother-in-law’s tongue leaf-section cuttings. Preformed Plantlets – A small number of house­ plants produce preformed plantlets along aerial shoots. Examples include spider plant, wax plant (Hoya) and strawberry begonia (Saxifraga stolonifera). Propagation is simple involving the removal of these plantlets (with or without aerial rootlets) and then inserting the base of the plantlets in a high-quality potting soil and treating the Figure 9. African violet leaf-petiole cuttings. plantlet as if it were a stem cutting.
  • 5. Outdoor Plants Many of your favorite outdoor shrubs, vines and ground covers can be easily propagated by cuttings. Softwood Cuttings Softwood cuttings are prepared from the soft, succulent, new flush of growth. Many ornamental plants can be propagated by softwood cuttings includ­ ing lilacs, forsythia, magnolia, weigela, oleander, pyracantha, spirea, apple, peach, pear and plum. While softwood cuttings tend to root easily and faster, they may require more care since the soft tissue is more susceptible to drying. In general, with softwood Figure 12. Hardwood cutting. cuttings, we are talking about cuttings that are typically 3 to 4 inches long, with two to three nodes and the basal leaves removed before inserting the Root Cuttings cutting into the media. Root cuttings are used very rarely and only on a few species that typically do not root well from stem Semi-Hardwood Cuttings cuttings. These include Aesculus, Aralia, Realistically, terms such as softwood and semi- Koelreuteria, sumac (Rhus) and sassafras. The most hardwood are not absolutes. Once the very soft flush challenging aspect of root cuttings is uncovering from of new growth starts to stiffen up and form “wood,” soil the roots of your desired plant and not some we have likely passed into the semi-hardwood stage. other plant. While root cuttings may be made at any A majority of our common ornamental shrubs are time of the year, it is probably more common to do propagated from semi-hardwood cuttings. Plants such this in the dormant season. Root pieces can be held in as viburnum, camellia, azalea, holly, crape myrtle, moist media in a cooler (>38 degrees), placed nearly hydrangea and euonymus use this type of cutting. horizontal in rooting media like you would treat other In Arkansas this peak time is often between May 15 cuttings or direct planted in well-prepared garden and June 15. soil with the top of the root-piece just barely emerging from the soil. Grafting/Budding Realistically, grafting or budding is not an easy technique for home gardeners to master. Most people would be amazed at how many ornamental plants in their landscape originated from budded or grafted plants. Many shade trees, all Japanese maples, many weeping plants and all hybrid tea roses use these techniques. Budding or grafting is also the mainstay of the tree fruit industry. Budding and grafting are essentially the same technique; however, with budding you are simply attaching the bud from your desired plant to the rootstock/understock, and with Figure 11. Dipping prepared holly cutting in rooting grafting you are using a piece of stem that contains hormone. many buds on it. The obvious advantage of bud­ ding is you can potentially get many more plants Hardwood Cuttings from your desired scion/stockplant. The major Hardwood cuttings are made from woody, dormant wood on deciduous plants (e.g., grape, poplar, rose, willow, peach, wisteria, elderberry, blackberry, forsythia) or some needle evergreens (e.g., junipers, arborvitae). Hardwood cuttings are easy to prepare and less perishable. For deciduous plants the cuttings are leafless and tend to be longer than soft­ wood cuttings. Hardwood cuttings are typically 6 to 12 inches long. As a general rule for the needle ever­ greens, rooting percentages tend to be increased when cuttings are harvested in mid- to late winter. As a general rule, higher rooting percentages occur on junipers when a small “heal” of wood is included at the base of the cutting. Figure 13. Chip budding.
  • 6. disadvantage of budding is that you are typically for plants such as wisteria, brambles, forsythia, working with a very small bud versus a larger piece weigela and spirea. Ground layering simply involves of stem that is easier to handle. A good propagation bending an intact shoot over and burying the middle text can go into the specific details related to budding portion (i.e., shoot tip comes back up above ground and grafting. and is exposed) under the existing soil. You may wish to wound the lower portion of the bent stem. With Layering easy-to-root plants, roots will often form along the Layering is a method of rooting a new plant while buried stem. Once roots have formed, you simply cut the stem is still attached to the parent plant. It is the stem below the rooted zone, excising the plant simple to perform and can be done in the home from the mother plant. without special equipment or structures. Division Air Layering – In the Many home gardeners did not realize that they case of air layering, we are were actually plant propagators when they divided forming roots on stems well common perennials such as cannas, hosta, daylilies, above the soil. This method iris, peonies, bleeding heart, many ornamental can be used successfully on a grasses, ferns and lily-of-the-valley. As a general rule, number of common tropical the best time to divide plants is during the dormant houseplants including croton, season. dumb cane (Dieffenbachia), dracaena, rubber plant and schefflera. Resources The process is quite Bubel, Nancy. 1988. The New Seed Starter’s simple. First, pick a point in Handbook. ISBN-10:0878577521. Figure 14. Air layering the stem at least 10 to 12 Mahonia. Dirr, M.A., and C.W. Heuser, J. 2006. The Reference inches from the tip. Remove Manual of Woody Plant Propagation: From Seed all leaves several inches on either side of the region to Tissue Culture: A Practical Working Guide to where you will make your air layer. The area will the Propagation of Over 1,100 Species. Varsity typically be about 6 inches in length. Two types of Press. ISBN-10:0942375092. Basically an A to Z stem cuts are typically used. One involves making an recipe book for propagating woody ornamental upward-angled cut into the stem that stops about plants. halfway through the stem. The other method involves Druse, Ken. 2000. Making More Plants: The Science, removing a ring of the outer bark. The ring of bark Art and Joy of Propagation. $32. ISBN-10: tissue will be 1 to 1.5 inches long. Apply a rooting 051770787X. hormone to the cut surfaces. If an angled or vertical Gowdy, Mary Ann. 2002. Home Propagation of House­ cut has been made into the stem, you will want to plants. University of Missouri - Columbia. G6560. prevent that from closing by inserting a small piece http://extension.missouri.edu/explorepdf/ of toothpick in the wounded area. Next, take a hand­ agguides/hort/g06560.pdf. ful of moist sphagnum moss and pack the moss in Hartmann, H.T., D.E. Kester, F.T. Davies, Jr., and and around the cut surfaces. To retain the moisture R.L. Geneve. 2001. Plant Propagation Principles around the wounded area for weeks/months, wrap a and Practices. Prentice-Hall. 7th edition. ISBN­ layer of plastic sheet around the entire work area 10:0136792359. This is considered the bible of and seal both ends with electrical tape. Many people plant propagation. It is used in every college and also cover the plastic with aluminum foil to prevent university. Expensive, but thorough. any light from penetrating the rooting zone and to Heffernan, Maureen. 1997. Burpee Seed Starter. John reflect any heat from sunlight. You can simply keep Wiley & Sons. ISBN-10:0028610024. your foliage plant in the same place (i.e., indoors or outdoors) where it was originally growing. Once roots do form, simply make a single cross-cut in the stem Supplies below the rooting zone. You should end up with roots • Your local garden supply store can often provide on the bottom and leaves at the top: a new plant. trays, media and rooting hormone. Ground Layering – Ground layering can be • A.M. Leonard, Piqua, OH, http://amleo.com/ used to easily propagate a number of ornamental • Hummert International, Earth City, MO, shrubs and vines. This simple technique works well http://www.hummert.com/ No endorsement is implied or discrimination intended for firms or references included or excluded from this publication. Printed by University of Arkansas Cooperative Extension Service Printing Services. DR. JAMES A. ROBBINS is professor and horticulture specialist - Issued in furtherance of Cooperative Extension work, Acts of May 8 ornamentals, University of Arkansas Division of Agriculture, and June 30, 1914, in cooperation with the U.S. Department of Cooperative Extension Service, Little Rock. Agriculture, Director, Cooperative Extension Service, University of Arkansas. The Arkansas Cooperative Extension Service offers its programs to all eligible persons regardless of race, color, national origin, religion, gender, age, disability, marital or veteran status, or any other legally protected status, and is an Affirmative FSA6024-PD-4-08RV Action/Equal Opportunity Employer.