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SPECIAL SENSES
The Senses
 General senses of touch (tactile)
 Temperature- thermoreceptors (heat)
 Pressure- mechanoreceptors (movement)
 Pain- mechanoreceptors
 Special senses
 Smell- chemoreceptors (chemicals)
 Taste- chemoreceptors
 Sight- photoreceptors (light)
 Hearing- mechanoreceptors
 Equilibrium- (balance) mechanoreceptors
2
The Eye and Vision
 70 percent of all sensory receptors are in the eyes
 Each eye has over a million nerve fibers
 Protection for the eye
 Most of the eye is enclosed in a bony orbit
made up of the lacrimal (medial), ethmoid
(posterior), sphenoid (lateral), frontal
(superior), and zygomatic and maxilla (inferior)
 A cushion of fat surrounds most of the eye
3
Accessory Structures of the Eye
 Eyelids- brush particles
out of eye or cover eye
 Eyelashes- trap
particles and keep them
out of the eye
4
Accessory Structures of the Eye
 Ciliary glands –
modified
sweat glands
between the
eyelashes- secrete acidic
sweat to kill bacteria,
lubricate eyelashes
5
Accessory Structures of the Eye
 Conjunctiva
 Membrane that
lines the eyelids
 Connects to the
surface of the
eye- forms a seal
 Secretes mucus
to lubricate the
eye
6
CONJUNCTIVITIS
- Inflammation of the conjunctiva
- Caused by bacterial or viral infection
- Highly contagious
7
Accessory Structures of the Eye
 Lacrimal
apparatus
 Lacrimal gland –
produces
lacrimal fluid
 Lacrimal canals
– drains lacrimal
fluid from eyes
8
Accessory Structures of the Eye
 Lacrimal sac –
provides passage of
lacrimal fluid
towards nasal cavity
9
Accessory Structures of the Eye
 Nasolacrimal
duct – empties
lacrimal fluid
into the nasal
cavity
10
Function of the Lacrimal Apparatus
 Properties of lacrimal fluid
 Dilute salt solution (tears)
 Contains antibodies (fight antigens- foreign
substance) and lysozyme (enzyme that destroys
bacteria)
 Protects, moistens, and lubricates the eye
 Empties into the nasal cavity
11
Extrinsic Eye Muscles
 Muscles attach to the outer surface of the eye
 Produce eye movements
12
When Extrinsic Eye Muscles Contract
 Superior oblique- eyes look out
and down
 Superior rectus- eyes looks up
 Lateral rectus- eyes look outward
 Medial rectus- eyes look inward
 Inferior rectus- eyes looks down
 Inferior oblique- eyes look in and
up
13
14
Structure of the Eye
15
 The wall is composed of three
tunics
 Fibrous tunic – outside layer
 Choroid – middle layer
 Sensory tunic – inside layer
The Fibrous Tunic
 Sclera
 White connective tissue layer
 Seen anteriorly as the “white of the eye”
 Semi-transparent
16
The Fibrous Tunic
 Cornea
 Transparent, central anterior portion
 Allows for light to pass through (refracts, or bends, light slightly)
 Repairs itself easily
 The only human tissue that can be transplanted without fear of
rejection
17
http://www.phys.ufl.edu/~avery/course/3400/vision/eye_photo.jpg
18
Choroid Layer
 Blood-rich nutritive tunic
 Pigment prevents light from scattering (opaque-
blocks light from getting in, has melanin)
19
Choroid Layer
 Modified interiorly into two structures
 Cilliary body – smooth muscle (contracts to adjust the shape of
the lens)
 Iris- pigmented layer that gives eye color (contracts to adjust
the size of the pupil- regulates entry of light into the eye)
 Pupil – rounded opening in the iris
20
Sensory Tunic (Retina)
 Contains receptor cells
(photoreceptors)
 Rods
 Cones
 Signals leave the
retina toward the
brain through the
optic nerve
21
Sensory Tunic (Retina)
 Signals pass from photoreceptors via a two-neuron chain
 Bipolar neurons and Ganglion cells
22
http://www.uams.edu/jei/patients/retina_services/images/retina.jpg
23
VISUAL PIGMENTS
Rhodopsin- visual purple, in high concentration in RODS
-Composed of opsin and retinal (a derivative of vitamin
A) proteins
-When light hits the protein it “bleaches”- turns yellow
and then colorless. It straightens out and breaks down
into opsin and retinal.
There are three different other opsins beside rhodopsin,
with absorption for yellowish-green (photopsin I), green
(photopsin II), and bluish-violet (photopsin III) light.
24
Neurons of the Retina and Vision
 Rods
 Most are found towards the edges of the retina
 Allow dim light vision and peripheral vision
(more sensitive to light, do not respond in bright
light)
 Perception is all in gray tones
25
ROD CELLS
26
Neurons of the Retina and Vision
 Cones
 Allow for detailed color vision
 Densest in the center of the retina
 Fovea centralis – area of the retina with only cones
 Respond best in bright light
 No photoreceptor cells are at the optic disk, or
blind spot
27
Cone Sensitivity
28
 There are three types of
cones
 Different cones are sensitive
to different wavelengths
- red- long - green-
medium - blue- short
 Color blindness is the result
of lack of one or more cone
type
How do we see colors?
• To see any color, the brain must compare the input
from different kinds of cone cells—and then make
many other comparisons as well.
• The lightning-fast work of judging a color begins in the
retina, which has three layers of cells. Signals from the
red and green cones in the first layer are compared by
specialized red-green "opponent" cells in the second
layer. These opponent cells compute the balance
between red and green light coming from a particular
part of the visual field. Other opponent cells then
compare signals from blue cones with the combined
signals from red and green cones. 29
COLORBLINDNESS
- An inherited trait that is
transferred on the sex
chromosomes (23rd pair)- sex-
linked trait
- Occurs more often in males
- Can not be cured or
corrected
•Comes from a lack of one or
more types of color receptors.
•Most are green or red or both
and that is due to a lack of red
receptors.
•Another possibility is to have
the color receptors missing
entirely, which would result in
black and white vision.
COLORBLINDNESS TEST PLATES
32
Lens
 Biconvex crystal-
like structure
 Held in place by
a suspensory
ligament
attached to the
ciliary body
 Refracts light
greatly
Internal Eye Chamber Fluids
 Aqueous humor
 Watery fluid found in chamber between
the lens and cornea
 Similar to blood plasma
 Helps maintain intraocular pressure
 Provides nutrients for the lens and cornea
 Reabsorbed into venous blood through
the canal of Schlemm
Refracts light
slightly
33
Internal Eye Chamber Fluids
 Vitreous humor
 Gel-like substance behind the lens
 Keeps the eye from collapsing
 Lasts a lifetime and is not replaced
http://faculty.washington.edu/kepeter/119/images/eye3.jpg
Refracts light
slightly
Holds lens
and retina in
place
34
Lens Accommodation
 Light must be focused to a point on
the retina for optimal vision
 The eye is set for distance vision
(over 20 ft away)
 20/20 vision- at 20 feet, you see what
a normal eye would see at 20 feet
(20/100- at 20, normal person would
see at 100)
 The lens must change shape to focus
for closer objects
35
Nearsightedness, or myopia is the
difficulty of seeing objects at a
distance.
Myopia occurs when the
eyeball is slightly longer than
usual from front to back. This
causes light rays to focus at a
point in front of the retina,
rather than directly on its
surface.
Concave lenses are used to
correct the problem.
MYOPIA
36
Hyperopia, or
farsightedness, is when
light entering the eye
focuses behind the retina.
Hyperoptic eyes are
shorter than normal.
Hyperopia is treated using
a convex lens.
http://web.mountain.net/~topeye/images/hyperopia.jpg
HYPEROPIA
37
Images Formed on the Retina
If the image is focused at the spot where the
optic disk is located, nothing will be seen.
This is known as the blind spot. There are
no photoreceptors there, as nerves and
blood vessels pass through this point.
38
Visual Pathway
 Photoreceptors of
the retina
 Optic nerve
 Optic nerve
crosses at the optic
chiasma
39
Visual Pathway
 Optic tracts
 Thalamus (axons
form optic
radiation)
 Visual cortex of the
occipital lobe
40
Eye Reflexes
 Internal muscles are controlled by the autonomic nervous
system
 Bright light causes pupils to constrict through action of
radial (iris) and ciliary muscles
 Viewing close objects causes accommodation
 External muscles control eye movement to follow objects-
voluntary, controlled at the frontal eye field
 Viewing close objects causes convergence (eyes moving
medially)
41
The Ear
 Houses two senses
 Hearing (interpreted in the auditory cortex of the
temporal lobe)
 Equilibrium (balance) (interpreted in the
cerebellum)
 Receptors are mechanoreceptors
 Different organs house receptors for each
sense
42
Anatomy of the Ear
 The ear is divided into three areas
 Outer (external) ear
 Middle ear
 Inner ear
43
The External Ear
 Involved in hearing only
 Structures of the external
ear
 Pinna (auricle)- collects
sound
 External auditory canal-
channels sound inward
44
The External Auditory Canal
 Narrow chamber in the temporal bone- through
the external auditory meatus
 Lined with skin
 Ceruminous (wax) glands are present
 Ends at the tympanic membrane (eardrum)
45
The Middle Ear or Tympanic Cavity
 Air-filled cavity within the temporal
bone
 Only involved in the sense of hearing
46
The Middle Ear or Tympanic Cavity
 Two tubes are associated with the inner ear
 The opening from the auditory canal is covered by the
tympanic membrane (eardrum)
 The auditory tube connecting the middle ear with the
throat (also know as the eustacian tube)
 Allows for equalizing pressure during yawning or
swallowing
 This tube is otherwise collapsed
47
Bones of the Tympanic Cavity
 Three bones span the
cavity
 Malleus (hammer)
 Incus (anvil)
 Stapes (stirrip)
48
Bones of the Tympanic Cavity
 Vibrations from eardrum move the
malleus
 These bones transfer sound to the
inner ear
49
50
Inner Ear or Bony Labyrinth
 Also known as osseous
labyrinth- twisted bony
tubes
 Includes sense organs
for hearing and balance
 Filled with perilymph
Inner Ear or Bony Labyrinth
Vibrations of the stapes push and pull on the membranous
oval window, moving the perilymph through the cochlea.
The round window is a membrane at the opposite end to
relieve pressure.
51
Inner Ear or Bony Labryinth
 A maze of bony chambers within the
temporal bone
 Cochlea
 Upper chamber is the scala
vestibuli
 Lower chamber is the scala
tympani
 Vestibule
 Semicircular
canals 52
Inner Ear or Bony Labyrinth
 Also known as osseous labyrinth-
twisted bony tubes
 Includes sense organs
for hearing and balance
 Filled with perilymph
Inner Ear or Bony Labyrinth
Vibrations of the stapes push and pull
on the membranous oval window, moving
the perilymph through the cochlea. The round window is
a membrane at the opposite end to relieve pressure.
Inner Ear or Bony Labryinth
 A maze of bony chambers within the
temporal bone
 Cochlea
 Upper chamber
is the scala
vestibuli
 Lower chamber
is the scala
tympani
 Vestibule
 Semicircular
canals
Organ of Corti
 Located within the cochlea
 Receptors = hair cells on the basilar
membrane
Scala tympani
Scala vestibuli
 Gel-like tectorial membrane is capable of
bending hair cells (endolymph in the
membranous labyrinth of the cochlear
duct flows over it and pushes on the
membrane)
Organ of Corti
Organs of Hearing
 Organ of Corti
 Cochlear nerve attached to hair cells
transmits nerve impulses to auditory cortex
on temporal lobe
Scala tympani
Scala vestibuli
Mechanisms of Hearing
 Vibrations from sound waves move
tectorial membrane (pass through the
endolymph fluid filling the
membranous labyrinth in the cochlear
duct)
 Hair cells are bent by the membrane
Mechanisms of Hearing
 An action potential starts in
the cochlear nerve
 The signal is transmitted to
the midbrain (for auditory
reflexes and then directed
to the auditory cortex of
the temporal lobe)
Continued stimulation can lead
to adaptation (over
stimulation to the brain
makes it stop interpreting
the sounds)
Mechanisms of Hearing
Organs of Equilibrium
 Receptor cells are in two structures
 Vestibule
 Semicircular canals
Organs of Equilibrium
 Equilibrium has two functional parts
 Static equilibrium- in the vestibule
 Dynamic equilibrium- in the semicircular
canals
Static Equilibrium
 Maculae –
receptors in
the
vestibule
 Report on
the
position of
the head
 Send
information
via the
vestibular
nerve
Static Equilibrium
 Anatomy of the maculae
 Hair cells are embedded in
the otolithic membrane
 Otoliths (tiny stones) float in
a gel around the hair cells
Function of Maculae
Movements cause otoliths to bend the
hair cells (gravity moves the “rocks”
over and pulls the hairs)
http://neuromedia.neurobio.ucla.edu/campbell/eyeandear/wp_images/177_macula_HP.gif
Dynamic Equilibrium
 Whole structure is the
ampulla
 Crista ampullaris –
receptors in the
semicircular canals
 Tuft of hair cells
 Cupula (gelatinous cap)
covers the hair cells
Dynamic Equilibrium
 Action of angular head
movements
 The cupula stimulates the hair
cells
 Movement of endolymph
pushes the
cupula over
and pulls the
hairs
 An impulse is
sent via the
vestibular nerve
to the cerebellum
DYNAMIC EQUILIBRIUM
STRUCTURES
http://www.faculty.une.edu/com/abell/histo/CristaAmp.jpg
Chemical Senses – Taste and
Smell
 Both senses use chemoreceptors
 Stimulated by chemicals in solution
 Taste has four types of receptors
 Smell can differentiate a large range of
chemicals
 Both senses complement each other
and respond to many of the same
stimuli
Olfaction – The Sense of Smell
 Olfactory receptors are in the roof of the nasal
cavity
 Neurons with long cilia
 Chemicals must be dissolved in mucus for
detection
Olfaction – The Sense of Smell
 Impulses are transmitted via the olfactory nerve
 Interpretation of smells is made in the cortex
(olfactory area of temporal lobe)
The Sense of Taste
 Taste buds
house the
receptor
organs
 Location of
taste buds
 Most are on
the tongue
 Soft palate
 Cheeks
The Tongue and Taste
 The tongue is covered with
projections called papillae
 Filiform papillae – sharp with no
taste buds
 Fungifiorm papillae – rounded
with taste buds
 Circumvallate papillae – large
papillae with taste buds
 Taste buds are found on the sides
of papillae
Structure of Taste Buds
 Gustatory cells are the receptors
 Have gustatory hairs (long microvilli)
 Hairs are stimulated by chemicals
dissolved in saliva
Structure of Taste Buds
 Impulses are carried to
the gustatory complex
(pareital lobe) by
several cranial nerves
because taste buds
are found in different
areas
 Facial nerve
 Glossopharyngeal
nerve
 Vagus nerve
Taste Sensations
 Sweet receptors
 Sugars
 Saccharine
 Some amino acids
 Sour receptors
 Acids
 Bitter receptors
 Alkaloids
 Salty receptors
 Metal ions
 Umami Glutamate, aspartate
(MSG, meats)
http://instruct1.cit.cornell.edu/courses/psych431/student2000/mle6/tonguebig.gif
Developmental Aspects of the
Special Senses
 Formed early in embryonic
development
 Eyes are outgrowths of the brain
 All special senses are functional at
birth
80

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Special senses

  • 2. The Senses  General senses of touch (tactile)  Temperature- thermoreceptors (heat)  Pressure- mechanoreceptors (movement)  Pain- mechanoreceptors  Special senses  Smell- chemoreceptors (chemicals)  Taste- chemoreceptors  Sight- photoreceptors (light)  Hearing- mechanoreceptors  Equilibrium- (balance) mechanoreceptors 2
  • 3. The Eye and Vision  70 percent of all sensory receptors are in the eyes  Each eye has over a million nerve fibers  Protection for the eye  Most of the eye is enclosed in a bony orbit made up of the lacrimal (medial), ethmoid (posterior), sphenoid (lateral), frontal (superior), and zygomatic and maxilla (inferior)  A cushion of fat surrounds most of the eye 3
  • 4. Accessory Structures of the Eye  Eyelids- brush particles out of eye or cover eye  Eyelashes- trap particles and keep them out of the eye 4
  • 5. Accessory Structures of the Eye  Ciliary glands – modified sweat glands between the eyelashes- secrete acidic sweat to kill bacteria, lubricate eyelashes 5
  • 6. Accessory Structures of the Eye  Conjunctiva  Membrane that lines the eyelids  Connects to the surface of the eye- forms a seal  Secretes mucus to lubricate the eye 6
  • 7. CONJUNCTIVITIS - Inflammation of the conjunctiva - Caused by bacterial or viral infection - Highly contagious 7
  • 8. Accessory Structures of the Eye  Lacrimal apparatus  Lacrimal gland – produces lacrimal fluid  Lacrimal canals – drains lacrimal fluid from eyes 8
  • 9. Accessory Structures of the Eye  Lacrimal sac – provides passage of lacrimal fluid towards nasal cavity 9
  • 10. Accessory Structures of the Eye  Nasolacrimal duct – empties lacrimal fluid into the nasal cavity 10
  • 11. Function of the Lacrimal Apparatus  Properties of lacrimal fluid  Dilute salt solution (tears)  Contains antibodies (fight antigens- foreign substance) and lysozyme (enzyme that destroys bacteria)  Protects, moistens, and lubricates the eye  Empties into the nasal cavity 11
  • 12. Extrinsic Eye Muscles  Muscles attach to the outer surface of the eye  Produce eye movements 12
  • 13. When Extrinsic Eye Muscles Contract  Superior oblique- eyes look out and down  Superior rectus- eyes looks up  Lateral rectus- eyes look outward  Medial rectus- eyes look inward  Inferior rectus- eyes looks down  Inferior oblique- eyes look in and up 13
  • 14. 14
  • 15. Structure of the Eye 15  The wall is composed of three tunics  Fibrous tunic – outside layer  Choroid – middle layer  Sensory tunic – inside layer
  • 16. The Fibrous Tunic  Sclera  White connective tissue layer  Seen anteriorly as the “white of the eye”  Semi-transparent 16
  • 17. The Fibrous Tunic  Cornea  Transparent, central anterior portion  Allows for light to pass through (refracts, or bends, light slightly)  Repairs itself easily  The only human tissue that can be transplanted without fear of rejection 17
  • 19. Choroid Layer  Blood-rich nutritive tunic  Pigment prevents light from scattering (opaque- blocks light from getting in, has melanin) 19
  • 20. Choroid Layer  Modified interiorly into two structures  Cilliary body – smooth muscle (contracts to adjust the shape of the lens)  Iris- pigmented layer that gives eye color (contracts to adjust the size of the pupil- regulates entry of light into the eye)  Pupil – rounded opening in the iris 20
  • 21. Sensory Tunic (Retina)  Contains receptor cells (photoreceptors)  Rods  Cones  Signals leave the retina toward the brain through the optic nerve 21
  • 22. Sensory Tunic (Retina)  Signals pass from photoreceptors via a two-neuron chain  Bipolar neurons and Ganglion cells 22
  • 24. VISUAL PIGMENTS Rhodopsin- visual purple, in high concentration in RODS -Composed of opsin and retinal (a derivative of vitamin A) proteins -When light hits the protein it “bleaches”- turns yellow and then colorless. It straightens out and breaks down into opsin and retinal. There are three different other opsins beside rhodopsin, with absorption for yellowish-green (photopsin I), green (photopsin II), and bluish-violet (photopsin III) light. 24
  • 25. Neurons of the Retina and Vision  Rods  Most are found towards the edges of the retina  Allow dim light vision and peripheral vision (more sensitive to light, do not respond in bright light)  Perception is all in gray tones 25
  • 27. Neurons of the Retina and Vision  Cones  Allow for detailed color vision  Densest in the center of the retina  Fovea centralis – area of the retina with only cones  Respond best in bright light  No photoreceptor cells are at the optic disk, or blind spot 27
  • 28. Cone Sensitivity 28  There are three types of cones  Different cones are sensitive to different wavelengths - red- long - green- medium - blue- short  Color blindness is the result of lack of one or more cone type
  • 29. How do we see colors? • To see any color, the brain must compare the input from different kinds of cone cells—and then make many other comparisons as well. • The lightning-fast work of judging a color begins in the retina, which has three layers of cells. Signals from the red and green cones in the first layer are compared by specialized red-green "opponent" cells in the second layer. These opponent cells compute the balance between red and green light coming from a particular part of the visual field. Other opponent cells then compare signals from blue cones with the combined signals from red and green cones. 29
  • 30. COLORBLINDNESS - An inherited trait that is transferred on the sex chromosomes (23rd pair)- sex- linked trait - Occurs more often in males - Can not be cured or corrected •Comes from a lack of one or more types of color receptors. •Most are green or red or both and that is due to a lack of red receptors. •Another possibility is to have the color receptors missing entirely, which would result in black and white vision.
  • 32. 32 Lens  Biconvex crystal- like structure  Held in place by a suspensory ligament attached to the ciliary body  Refracts light greatly
  • 33. Internal Eye Chamber Fluids  Aqueous humor  Watery fluid found in chamber between the lens and cornea  Similar to blood plasma  Helps maintain intraocular pressure  Provides nutrients for the lens and cornea  Reabsorbed into venous blood through the canal of Schlemm Refracts light slightly 33
  • 34. Internal Eye Chamber Fluids  Vitreous humor  Gel-like substance behind the lens  Keeps the eye from collapsing  Lasts a lifetime and is not replaced http://faculty.washington.edu/kepeter/119/images/eye3.jpg Refracts light slightly Holds lens and retina in place 34
  • 35. Lens Accommodation  Light must be focused to a point on the retina for optimal vision  The eye is set for distance vision (over 20 ft away)  20/20 vision- at 20 feet, you see what a normal eye would see at 20 feet (20/100- at 20, normal person would see at 100)  The lens must change shape to focus for closer objects 35
  • 36. Nearsightedness, or myopia is the difficulty of seeing objects at a distance. Myopia occurs when the eyeball is slightly longer than usual from front to back. This causes light rays to focus at a point in front of the retina, rather than directly on its surface. Concave lenses are used to correct the problem. MYOPIA 36
  • 37. Hyperopia, or farsightedness, is when light entering the eye focuses behind the retina. Hyperoptic eyes are shorter than normal. Hyperopia is treated using a convex lens. http://web.mountain.net/~topeye/images/hyperopia.jpg HYPEROPIA 37
  • 38. Images Formed on the Retina If the image is focused at the spot where the optic disk is located, nothing will be seen. This is known as the blind spot. There are no photoreceptors there, as nerves and blood vessels pass through this point. 38
  • 39. Visual Pathway  Photoreceptors of the retina  Optic nerve  Optic nerve crosses at the optic chiasma 39
  • 40. Visual Pathway  Optic tracts  Thalamus (axons form optic radiation)  Visual cortex of the occipital lobe 40
  • 41. Eye Reflexes  Internal muscles are controlled by the autonomic nervous system  Bright light causes pupils to constrict through action of radial (iris) and ciliary muscles  Viewing close objects causes accommodation  External muscles control eye movement to follow objects- voluntary, controlled at the frontal eye field  Viewing close objects causes convergence (eyes moving medially) 41
  • 42. The Ear  Houses two senses  Hearing (interpreted in the auditory cortex of the temporal lobe)  Equilibrium (balance) (interpreted in the cerebellum)  Receptors are mechanoreceptors  Different organs house receptors for each sense 42
  • 43. Anatomy of the Ear  The ear is divided into three areas  Outer (external) ear  Middle ear  Inner ear 43
  • 44. The External Ear  Involved in hearing only  Structures of the external ear  Pinna (auricle)- collects sound  External auditory canal- channels sound inward 44
  • 45. The External Auditory Canal  Narrow chamber in the temporal bone- through the external auditory meatus  Lined with skin  Ceruminous (wax) glands are present  Ends at the tympanic membrane (eardrum) 45
  • 46. The Middle Ear or Tympanic Cavity  Air-filled cavity within the temporal bone  Only involved in the sense of hearing 46
  • 47. The Middle Ear or Tympanic Cavity  Two tubes are associated with the inner ear  The opening from the auditory canal is covered by the tympanic membrane (eardrum)  The auditory tube connecting the middle ear with the throat (also know as the eustacian tube)  Allows for equalizing pressure during yawning or swallowing  This tube is otherwise collapsed 47
  • 48. Bones of the Tympanic Cavity  Three bones span the cavity  Malleus (hammer)  Incus (anvil)  Stapes (stirrip) 48
  • 49. Bones of the Tympanic Cavity  Vibrations from eardrum move the malleus  These bones transfer sound to the inner ear 49
  • 50. 50 Inner Ear or Bony Labyrinth  Also known as osseous labyrinth- twisted bony tubes  Includes sense organs for hearing and balance  Filled with perilymph
  • 51. Inner Ear or Bony Labyrinth Vibrations of the stapes push and pull on the membranous oval window, moving the perilymph through the cochlea. The round window is a membrane at the opposite end to relieve pressure. 51
  • 52. Inner Ear or Bony Labryinth  A maze of bony chambers within the temporal bone  Cochlea  Upper chamber is the scala vestibuli  Lower chamber is the scala tympani  Vestibule  Semicircular canals 52
  • 53. Inner Ear or Bony Labyrinth  Also known as osseous labyrinth- twisted bony tubes  Includes sense organs for hearing and balance  Filled with perilymph
  • 54. Inner Ear or Bony Labyrinth Vibrations of the stapes push and pull on the membranous oval window, moving the perilymph through the cochlea. The round window is a membrane at the opposite end to relieve pressure.
  • 55. Inner Ear or Bony Labryinth  A maze of bony chambers within the temporal bone  Cochlea  Upper chamber is the scala vestibuli  Lower chamber is the scala tympani  Vestibule  Semicircular canals
  • 56. Organ of Corti  Located within the cochlea  Receptors = hair cells on the basilar membrane Scala tympani Scala vestibuli
  • 57.  Gel-like tectorial membrane is capable of bending hair cells (endolymph in the membranous labyrinth of the cochlear duct flows over it and pushes on the membrane) Organ of Corti
  • 58. Organs of Hearing  Organ of Corti  Cochlear nerve attached to hair cells transmits nerve impulses to auditory cortex on temporal lobe Scala tympani Scala vestibuli
  • 59. Mechanisms of Hearing  Vibrations from sound waves move tectorial membrane (pass through the endolymph fluid filling the membranous labyrinth in the cochlear duct)  Hair cells are bent by the membrane
  • 60. Mechanisms of Hearing  An action potential starts in the cochlear nerve  The signal is transmitted to the midbrain (for auditory reflexes and then directed to the auditory cortex of the temporal lobe)
  • 61. Continued stimulation can lead to adaptation (over stimulation to the brain makes it stop interpreting the sounds) Mechanisms of Hearing
  • 62. Organs of Equilibrium  Receptor cells are in two structures  Vestibule  Semicircular canals
  • 63. Organs of Equilibrium  Equilibrium has two functional parts  Static equilibrium- in the vestibule  Dynamic equilibrium- in the semicircular canals
  • 64. Static Equilibrium  Maculae – receptors in the vestibule  Report on the position of the head  Send information via the vestibular nerve
  • 65. Static Equilibrium  Anatomy of the maculae  Hair cells are embedded in the otolithic membrane  Otoliths (tiny stones) float in a gel around the hair cells
  • 66. Function of Maculae Movements cause otoliths to bend the hair cells (gravity moves the “rocks” over and pulls the hairs)
  • 68. Dynamic Equilibrium  Whole structure is the ampulla  Crista ampullaris – receptors in the semicircular canals  Tuft of hair cells  Cupula (gelatinous cap) covers the hair cells
  • 69. Dynamic Equilibrium  Action of angular head movements  The cupula stimulates the hair cells  Movement of endolymph pushes the cupula over and pulls the hairs  An impulse is sent via the vestibular nerve to the cerebellum
  • 71. Chemical Senses – Taste and Smell  Both senses use chemoreceptors  Stimulated by chemicals in solution  Taste has four types of receptors  Smell can differentiate a large range of chemicals  Both senses complement each other and respond to many of the same stimuli
  • 72. Olfaction – The Sense of Smell  Olfactory receptors are in the roof of the nasal cavity  Neurons with long cilia  Chemicals must be dissolved in mucus for detection
  • 73. Olfaction – The Sense of Smell  Impulses are transmitted via the olfactory nerve  Interpretation of smells is made in the cortex (olfactory area of temporal lobe)
  • 74. The Sense of Taste  Taste buds house the receptor organs  Location of taste buds  Most are on the tongue  Soft palate  Cheeks
  • 75. The Tongue and Taste  The tongue is covered with projections called papillae  Filiform papillae – sharp with no taste buds  Fungifiorm papillae – rounded with taste buds  Circumvallate papillae – large papillae with taste buds  Taste buds are found on the sides of papillae
  • 76. Structure of Taste Buds  Gustatory cells are the receptors  Have gustatory hairs (long microvilli)  Hairs are stimulated by chemicals dissolved in saliva
  • 77. Structure of Taste Buds  Impulses are carried to the gustatory complex (pareital lobe) by several cranial nerves because taste buds are found in different areas  Facial nerve  Glossopharyngeal nerve  Vagus nerve
  • 78. Taste Sensations  Sweet receptors  Sugars  Saccharine  Some amino acids  Sour receptors  Acids  Bitter receptors  Alkaloids  Salty receptors  Metal ions  Umami Glutamate, aspartate (MSG, meats) http://instruct1.cit.cornell.edu/courses/psych431/student2000/mle6/tonguebig.gif
  • 79. Developmental Aspects of the Special Senses  Formed early in embryonic development  Eyes are outgrowths of the brain  All special senses are functional at birth
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