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(8.5095°N, 81.1173°W), 900 m elev. 2 August 2008. Leonhard Stadler and Nadim Hamad. SMF 89577. This record extends by 200 m, the upper confirmed elevational limit for the species previously reported by Savage (2002) and Köhler (2008). The presence of L. southi around Alto de Piedra was documented by Martínez and Rodriguez (1992). However, elevational distribu- tion for their individual records was unspecified; only a general elevational range of between 800 m and 1450 m was given. Hoplocercidae Enyalioides heterolepis (Bocourt’s Dwarf Iguana). VERAGUAS: Cerro Negro, ca. 6 km NNW of Santa Fé (8.5690°N, 81.0989°W), 700 m elev. 30 July 2008. Sebastian Lotzkat. SMF 89558. Cerro Negro, 6 km NNW of Santa Fé (8.5693°N, 81.1022°W), 713 m fig. 1. Generalized localities in western Panamá visited by the authors elev. 28 June 2008. Arcadio Carrizo. SMF 89955–56. First re-in 2008: 1) Jurutungo, Chiriquí; 2) Reserva Forestal La Fortuna, Chiriquí; cords for Veraguas, extending the range more than 50 km W3) road to Cerro Colorado, Comarca Ngöbe-Buglé; 4) La Nevera, of the formerly westernmost record from El Copé, Coclé (LipsComarca Ngöbe-Buglé; 5) Cerro Mariposa (= Cerro Tute), Veraguas; 6)Cerro Negro, Veraguas; 7) Río Chilagres, Veraguas. 1999). Köhler (2008) mentioned a record of E. heterolepis from Bocas del Toro, which as far as we can tell is not supported by a voucher specimen. A now lost individual from Cerro Naricesthose 17 species, 11 of which are new provincial records; other (between Cerro Negro and Río Chilagres, see Fig. 1) that wasaccounts list new elevational parameters for the species. listed as Sceloporus squamosus by Martínez et al. (1994), actu- Abbreviations for museum collections follow those of Leviton ally represented an E. heterolepis (A. Rodriguez, pers. comm.).et al. (1985). Elevations were rounded to the next tenth. The map(Fig. 1) was created using DIVA-GIS and the NASA elevation Polychrotidaedatasets processed by Jarvis et al. (2006); map datum is WGS84. Anolis auratus (Grass Anole). VERAGUAS: road between Santi-All vouchers were verified by Javier Sunyer. Common names ago and Santa Fé, ca. 8 km N of Santa Fé; (8.1796°N, 80.951°W),came from Frank and Ramus (1995), except for Leposoma southi 80 m elev. 14 May 2008. Andreas Hertz. SMF 89444. First record(Uetz 2008), Urotheca guentheri (Solórzano 2004), and by us for Veraguas, bridging the gap between localities in Chiriquí toherein for Anolis fortunensis and A. pseudopachypus, two species the west and Los Santos, Colón, and Panamá to the east (Köhlerfor which there are no published common names. We decided et al. 2008; Young et al. 1999).that for those two species, the common names should denote theirtype localities; Reserva Forestal La Fortuna, Panamá, and La Ne- Anolis capito (Bighead Anole). CHIRIQUÍ: Reserva Forestal Lavera, Panamá, respectively. Fortuna (8.6744°N, 82.2161°W), 1460 m elev. 16 June 2008. An- dreas Hertz. SMF 89449. This record increases the known eleva- Squamata — Lizards tional limit for the species by 160 m (Köhler 2008). Sphaerodactylidae Anolis fortunensis (La Fortuna Anole). CHIRIQUÍ: Reserva For-Lepidoblepharis xanthostigma (Costa Rica Scaly-eyed Gecko). estal La Fortuna (8.6776°N, 82.1981°W), 1750 m elev. 27 JuneVERAGUAS: Cerro Mariposa, near Alto de Piedra, ca. 3.5 km W 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89466. Thisof Santa Fé (8.5161°N, 81.1185°W), 880 m elev. 12 May 2008. record extends the known elevational range of the species by 550Andreas Hertz and Sebastian Lotzkat. SMF 89576. Cerro Negro, m (Köhler 2008).ca. 6 km NNW of Santa Fé (8.5690°N, 81.0989°W), 700 m elev. Anolis humilis (Humble Anole). CHIRIQUÍ: Reserva Forestal28 July 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89963. La Fortuna (8.6761°N, 82.2006°W), 1660 m elev. 27 June 2008.Martínez and Rodriguez (1992), in their inventory of “Cerro Sebastian Lotzkat. SMF 89480. This record increases the knownTute” (= Cerro Mariposa) and Martínez et al. (1994) in their elevational range by 60 m (Köhler 2008).species list of the nearby mountains, Narices and La Anselma, Anolis pseudopachypus (La Nevera Anole). CHIRIQUÍ: Reser-both reported the presence of Lepidoblepharis sp., and consid- va Forestal La Fortuna (8.6761°N, 82.2006°W), 1660 m elev.;ered them to resemble L. xanthostigma. However, because the (8.6793°N, 82.193°W), 1800 m elev. 25 and 27 June 2008. Andre-specific identities of these geckos were not made and the respec- as Hertz and Sebastian Lotzkat. SMF 89523, 89526, respectively.tive voucher specimens were lost (V. Martínez pers. comm.), our First records for Chiriquí, and extends the range ca. 50 km WNWrecords represent the first confirmed occurrence for the species of the type locality (La Nevera, Comarca Ngöbe-Buglé; Köhlerin Veraguas. The records also fill the gap between records to the et al. 2007). COMARCA NGÖBE-BUGLÈ: Valley north of Lawest in Chiriquí and Bocas del Toro and the east in Coclé and Nevera, ca. 5.5 km N of Hato Chamí (8.5018°N, 81.7689°W),Colón (Young et al. 1999). 1560 m elev. 9 August 2008. Andreas Hertz and Sebastian Lotz- Gymnophthalmidae kat. SMF 89519–89521. Ridgetop NE of La Nevera, ca. 5.5 km N of Hato Chami (8.4954°N, 81.7673°W), 1810 m elev. 18 AugustLeposoma southi (Northern Spectacled Lizard). VERAGUAS: 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89522. TheseCerro Mariposa, near Alto de Piedra, ca. 3.5 km W of Santa Fé records, near the type locality of Anolis pseudopachypus, repre- Herpetological Review 41(4), 2010 521
sent the lowest and highest elevational records, respectively, for Viperidaethe species, which was previously known only from 1600 m el-evation (Köhler et al. 2007). Bothriechis nigroviridis (Black-speckled Palm Viper). COMAR- CA NGÖBE-BUGLÈ: La Nevera, ca. 5.5 km N of Hato ChamíAnolis salvini (Salvin’s Anole). CHIRIQUÍ: Jurutungo, near bor- (8.4996°N, 81.77°W), 1650 m elev. 11 May 2008. Andreas Hertz,der with Costa Rica (8.912°N, 82.7182°W), 1980 m elev. 8 July Leonhard Stadler, Nadim Hamad, and Sebastian Lotzkat. SMF2008. Andreas Hertz and Sebastian Lotzkat. SMF 89527. This 89534. First record for Comarca Ngöbe-Buglé, and an 80 kmrecord extends the known elevational distribution of the species E range extension from the formerly easternmost record in Bo-upward ca. 150 m (Köhler 2008; Savage 2002). quete, Chiriquí (Dunn 1947).Anolis vittigerus (Garland Anole). VERAGUAS: Río Chilagres, Bothriechis schlegelii (Eyelash Palm Viper). VERAGUAS: Cer-ca. 9 km NE of Santa Fé (8.5872°N, 81.0363°W), 480 m elev. 3 ro Mariposa, near Alto de Piedra, approx. 3.5 km W of SantaJune 2008. Andreas Hertz and Sebastian Lotzkat. SMF 89528. Fé (8.5157°N, 81.1197°W), 860 m elev. 13 May 2008. GuntherFirst record for Veraguas, extending the range ca. 50 km W of the Köhler. SMF 89535. First record for Veraguas, which fills the gapformerly westernmost record from north of El Copé, Coclé (Poe between Isla Escudo de Veraguas, Comarca Ngöbe-Buglé (ca. 85and Ibañez 2007). km NW), and El Valle de Anton, Coclé (ca. 105 km E) (Pérez- Squamata — Snakes Santos 1999). Colubridae Acknowledgments.—Collecting and exportation permits SE/A-30-08 and SEX/A-108-08 were provided by A. Salazar, Y. Hidalgo, and J. Gar-Dipsas articulata (American Snail-eater). VERAGUAS: Cerro cía, Autoridad Nacional del Ambiente (ANAM), Panamá City, Panamá.Negro, ca. 6 km NNW of Santa Fé (8.5756°N, 81.0976°W), 1000 Q. D. Fuenmayora and V. Martínez provided valuable assistance with ac-m elev. 29 July 2008. Arcadio Carrizo. SMF 89552. Since locali- quisition of permits. An additional permit for the Comarca Ngöbe-Bugléty data for the holotype of D. articulata is imprecise (see remarks was issued by A. Montezuma (ANAM), San Félix, Panamá. For fieldin Peters 1960), this is the first validated record for Veraguas. The assistance and logistical support, we thank C. O’Shea and S. Abrego, A. Bennett, P. Yangüez, O. Cáceres and family, M. Piepenbring, andrecord reconfirms the species’ occurrence at 1000 m, reported as the Peña Solís family. A. Carrizo received financial support from theits upper elevational limit by Solórzano (2004). DAAD. Field work funding to A. Hertz came from FAZIT-Stiftung andDipsas temporalis (Temporal Snail-eater). VERAGUAS: Cer- from Studienstiftung des deutschen Volkes to S. Lotzkat.ro Mariposa, near Alto de Piedra, ca. 3.5 km W of Santa Fé(8.5107°N, 81.1207°W), 990 m elev. 10 July 2008. Leonhard literature citedStadler and Nadim Hamad. SMF 89552. First record for Vera- cadle, J. e., and c. W. MYerS. 2003. Systematics of snakes referred toguas, and a range extension of ca.130 km W of the South slope Dipsas variegata Panama and western South America, with revalida-of Cerro La Campana, the formerly westernmost record in Pan- tion of two species and notes on defensive behaviors in the Dipsadiniamá (Pérez-Santos 1999). Another specimen (SMF 89553) found (Colubridae). Am. Mus. Novit. :1–47.nearby (8.5245°N, 81.1332°W), 700 m elev., on 5 August 2008 dunn, e. r. 1947. Snakes of the Lérida Farm (Chiriqui Volcano, westernwas acquired at the lowest elevation reported for this species Panamá). Copeia 1947:153–157.(Pérez-Santos 1999). frank, n., and e. raMuS. 1995. A Complete Guide to Scientific and Common Names of Reptiles and Amphibians of the World. N. G.Imantodes cenchoa (Blunt-headed Tree Snake). COMARCA Publ., Inc., Pottsville, Pennsylvania.NGÖBE-BUGLÈ: road to Cerro Colorado west of La Nevera, HuleBak, e., S. Poe, r. iBáñez, and e. e. WilliaMS. 2007. A strikingca. 10 km NNW of Hato Chamí (8.5303°N, 81.7987°W), 1660 m new species of Anolis lizard (Squamata, Iguania) from Panama. Phyl-elev. 10 May 2008. Andreas Hertz. SMF 89573. The locality is lomedusa 6:5–10.situated at an elevation 160 m above that reported for the species JarviS, a., H. i. reuter, a. nelSon, and e. guevara. 2006. Hole-filledby Solórzano (2004). seamless SRTM data V3, International Centre for Tropical Agriculture (CIAT), available from http://srtm.csi.cgiar.org. Downloaded Novem-Oxyrhopus petola (Calico False Coral Snake). VERAGUAS: Río ber 2007.Chilagres, ca. 9 km NE of Santa Fé (8.5872°N, 81.0363°W), 480 köHler, g. 2008. Reptiles of Central America, 2nd ed. Herpeton, Verlagm elev. 3 June 2008. Andreas Hertz and Sebastian Lotzkat. SMF Elke Köhler, Offenbach, Germany.89585. First record for Veraguas, which bridges a distributional –––––. 2010. A revision of the Central American species related to Ano-gap between localities in the provinces of Chiriquí and Bocas del lis pentaprion, with the resurrection of A. beckeri and the descriptionToro to the west and Colón and Panamá to the east (Young et al. of a new species (Squamata Polychrotidae). Zootaxa 2354:1–18. –––––, S. lotzkat, and a. Hertz. 2010. A new species of Sibon (Squa-1999). mata: Colubridae) from Western Panama. Herpetologica 66:80–85.Urotheca guentheri (Striped Glasstail). VERAGUAS: Cer- –––––, M. Ponce, J. SunYer, and a. BatiSta. 2007. Four new speciesro Mariposa, near Alto de Piedra, ca. 3.5 km W of Santa Fé of anoles (genus Anolis) from the Serranía de Tabasará, west-central(8.5002°N, 81.1175°W), 1250 m elev. 5 June 2008. Leonhard Panama (Squamata: Polychrotidae). Herpetologica 63:375–391.Stadler and Nadim Hamad. SMF 89603. First record for Vera- –––––, and J. SunYer. 2008. Two new species of anoles formerly refer- red to as Anolis limifrons (Squamata: Polychrotidae). Herpetologicaguas, filling a gap between localities in the provinces of Bocas 64:92–108.del Toro (La Loma, ca.120 km NW) and Coclé (about 110 km E) –––––, –––––, M. Ponce, and a. BatiSta. 2008. Noteworthy records of(Myers 1974). amphibians and reptiles in Panama (Amphibia: Plethodontidae, Crau-522 Herpetological Review 41(4), 2010