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September 6

What are tree rings?
Source: dylancantwell
MAJOR TOPICS
What are trees? of basic anatomy The cambium
            Tour
GEOG5839
 WHAT ARE TREES?
Tree at Chancellorsville




Source: Andy Frasse o
Trees are not limited to a single order or even division of plants.



          “Tree-ness” is defined by a specific

   GROWTH FORM.
Trees are perennial plants, which are plants
that live for more than two years.
Trees have a single main stem (or trunk).
Shrubs have multiple stems
(and are not trees).
Plants are o en required to a ain a minimum height at maturity
to qualify as trees (usually between 3 and 6 meters tall).
Trees are woody plants (a category that also includes shrubs
and some vines).
Woody plants are capable of secondary thickening,
adding new yearly growth layers of cells onto the
accumulated growth of previous years.
Most recent
               growth




Oldest wood
Cross-sectional plane
Cross-sectional
                                     (or transverse) plane



                                     Radial plane




                                    Tangential plane

Source: Hoadley, Identifying Wood
Photograph: Dave_Gray
GEOG5839
TOUR OF BASIC ANATOMY
Heartwood

              Sapwood
Cambium


               Bark
tissue containing plant ‘stem’ cells




The vascular cambium is a lateral meristem
in the vascular tissue of plants.
The vascular cambium is also the source of both
the secondary xylem (inwards, towards the pith)
and the secondary phloem (outwards), and is
located between these tissues in the stem and root.
Photograph: easagor
Bark
Pith




       Cambium
The pith is the small core of so spongey tissue
that forms the central longitudinal axis of a tree
stem, branch or twig.
Scots pine
                              Pinus sylvestris
Source: Fritz Schweingruber
Common beech
                              Fagus sylvestris
Source: Fritz Schweingruber
Sapwood
Source: sonykus
Heartwood

            Ph
              oto
                 syn
                    the
                       tic
                             wa
                               ste
                                  s
GYMNOSPERMS

  VS.
ANGIOSPERMS
Gymnosperms
 (coniferous)
Gymnosperms like this Douglas-fir have encapsulated seeds.
Red pine
Pinus resinosa
Earlywood


                           Latewood


                   Total ring-width




Eastern hemlock
Tsuga canadensis
Tracheids




Scots pine
Pinus sylvestris
Tracheids are elongated cells that function
in food conduction and structural support.
large cells
                                               thin cell walls

                                    small cells
                                  thick cell walls




        Scots pine
        Pinus sylvestris
Photograph: Fritz Schweingruber
Ray




Scots pine
Pinus sylvestris
Rays are fla ened bands of tissue that extend
horizontally in a radial plane through the tree stem.
Angiosperms
(deciduous)
White oak
Quercus alba
Angiosperms like this oak have exposed seeds.
Bur oak
Quercus macrocarpa
Earlywood vessels




English oak
Quercus robur
Latewood
                 vessels




English oak
Quercus robur
Vessel elements are a type of hardwood cell
that has a relatively large diameter, thick cell wall
and perforate (open) ends.
Fibre



                                          English oak
                                          Quercus robur
Photograph: Fritz Schweingruber
Fibres are an elongated hardwood cell with thick walls
that contributes greatly to the material strength of wood.
Ray


                                        English oak
                                        Quercus robur
Photograph: Fri s Schweingruber
RING POROUS   DIFFUSE POROUS
Bur oak
Quercus macrocarpa
Sycamore maple
Acer psudoplantanus
European beach
Fagus sylvatica
A growth-ring boundary in Chamaecyparis lawsoniana (Port-Orford cedar)




Source: Bu erfield and Meylan (1980)
GEOG5839
 THE CAMBIUM
tissue containing plant ‘stem’ cells




The vascular cambium is a lateral meristem
in the vascular tissue of plants.
ca
  m
 bi
  um



          phloem




  xylem
CHARACTERISTICS OF THE CAMBIUM



1
Self-sustaining
The cambium is a self-sustaining system, and retains its
functions for a long time (sometimes for centuries or millennia).
Bristlecone pine
          Pinus longaeva
Photograph: mlhradio
CHARACTERISTICS OF THE CAMBIUM



2   Resource-intensive
    As the cambium grows, it consumes scare resources that cannot
    be used for growth by the rest of the tree.
Limber pine
Pinus flexilis
Limber pine
Pinus flexilis
CHARACTERISTICS OF THE CAMBIUM



3   Flexible
    Cells produced by the cambium (called ‘cambial
    derivatives) can be differentiated into the range of cells
    found in the xylem and phloem.
Pinus spp.
Source: Marchand, P. J. 1996. Life in the Cold: An
Introduction to Winter Ecology, 3rd ed. University
Press of New England. Hanover, NH. 304 pp.
DORMANT CAMBIUM

      1 - 10 cells
  viscous protoplasm
   sharp cell walls
Source: Ben Cooper
ca
  m
 bi
  um



          phloem




  xylem
DORMANT CAMBIUM        ACTIVE CAMBIUM

      1 - 10 cells          8 - 20 cells
  viscous protoplasm      fluid protoplasm
   sharp cell walls    semi-transparent walls
CAMBIAL GROWTH
Radi al file




                                                Scots pine
                                                Pinus sylvestris
Photograph: Fritz Schweingruber
Latewood
               (prior year)
                                                   phloem

Early season

                              Immature earlywood
Latewood
             (prior year)
                                      Immature earlywood


Mid-season

                                                           phloem
                        Mature earlywood
Mature earlywood   Immature
                                 latewood

Late season
                                            phloem
A growth-ring boundary in Chamaecyparis lawsoniana (Port-Orford cedar)




Source: Bu erfield and Meylan (1980)
September 6

What are tree rings?
GEOG5839
  XT C L AS S
NE

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