Protein synthesis ribosome

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Dr Muhammad Mustansar

Protein Synthesis



Assembly is coupled w/ transcription and prerRNA processing

Prokaryotic
Translational
Initiation

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of Translation

Recycling of EF-Tu

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Active
enzyme

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Protein synthesis ribosome

Protein synthesis ribosome

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Most bacteria are free-living organisms that grow by increasing in mass and then divide by binary fission. Growth and division are controlled by genes, the expression of which must be regulated appropriately. Genes whose activity is controlled in response to the needs of a cell or organism are called regulated genes. All organisms also have a large number of genes whose products are essential to the normal functioning of a growing and dividing cell, no matter what the conditions are. These genes are always active in growing cells and are known as constitutive genes or housekeeping genes; examples include genes that code for the enzymes needed for protein synthesis and glucose metabolism. Note that all genes are regulated on some level. If normal cell function is impaired for some reason, the expression of all genes, including constitutive genes, is reduced by regulatory mechanisms. Thus, the distinction between regulated and constitutive genes is somewhat arbitrary.

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Transcription in eukaryotes: A brief view Transcription is the process by which single stranded RNA is synthesized by double stranded DNA. Transcription in eukaryotes and prokaryotes has many similarities while at the same time both showing their individual characteristics due to the differences in organization. RNA Polymerase (RNAP or RNA Pol) is different in prokaryotes and eukaryotes. Coupled transcription is seen in prokaryotes but not in Eukaryotes. In eukaryotes the pre-RNA should be spliced first to be translated. In Eukaryotic transcription, synthesis of RNA occurs in the 3’→5’ direction. The 3’ end is more reactive due to the hydroxide group. 5’ end containing phosphate groups meanwhile, is not very reactive when it comes to adding new nucleotides. In Eukaryotes, the whole genome is not transcribed at once. Only a part of the genome is transcribed which also acts as the first, principle stage of genetic regulation. Eukaryotes have five nuclear polymerases: • RNA Polymerase I: This produces rRNA (23S, 5.8S, and 18S) which are the major components in a ribosome. This also produces pre-rRNA in yeasts. • RNA Polymerase II: Helps in the production of mRNA (messenger RNA), snRNA (small, nuclear RNA), miRNA. This is the most studied type and requires several transcription factors for its binding • RNA Polymerase III: This synthesizes tRNA (transfer RNA), 5S rRNA and other small RNAs required in the cytosol and nucleus. • RNA Polymerase IV: Synthesizes siRNA (small interfering RNA) in plants. • RNA Polymerase V: This is the least studied polymerase and synthesizes siRNA-directed heterochromatin in plants. Eukaryotic transcription can be broadly divided into 4 stages: • Pre-Initiation • Initiation • Elongation • Termination Transcription is an elaborate process which cells use to copy the genetic information stored in DNA into RNA. This pre-RNA is modified into mRNA before being transcribed to proteins. Transcription is the first step to utilizing the genetic information in a cell. Both Eukaryotes and Prokaryotes employ this process with the basic phases remaining the same. However eukaryotic transcription is more complex indicating the changes transcription has undergone towards perfection during evolution.

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Protein synthesis ribosome

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2. Protein Synthesis

3.

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6.  Assembly is coupled w/ transcription and prerRNA processing

7.

8.

9. Prokaryotic Translational Initiation

10. Eukaryotic Initiation of Translation

11.

12.

13. Recycling of EF-Tu

14.

15.

16.

17. Regulation of Gene Expression RNA Processing 5’CAP Active enzyme Post-translational modification mRNA AAAAAA RNA Degradation Protein Degradation

18. Regulation of Globin gene translation by heme