A stage-structured delayed advection reaction-diffusion model for single species

I
β€’β€’

In this paper, we derived a delay advection reaction-diffusion equation with linear advection term from a stage-structured model, then the derived equation is used under the homogeneous Dirichlet boundary conditions ν‘’ ν‘š ( 0, ν‘‘ ) = 0, ν‘’ ν‘š ( 퐿, ν‘‘ ) = 0, and the initial condition ν‘’ ν‘š ( ν‘₯, 0 ) = ν‘’ ( ν‘₯ ) > 0, ν‘₯ ∈ [βˆ’νœ, 0] with ν‘’ ν‘š 0 (0) > 0 in order to find the minimum value of domain 퐿 that prevents extinction of the species under the effect of advection reaction-diffusion equation. Finally, for the measurement the time lengths from birth to the development of the species population, time delays are integrated. ν‘š 0

International Journal of Electrical and Computer Engineering (IJECE)
Vol. 10, No. 6, December 2020, pp. 6260~6267
ISSN: 2088-8708, DOI: 10.11591/ijece.v10i6.pp6260-6267  6260
Journal homepage: http://ijece.iaescore.com/index.php/IJECE
A stage-structured delayed advection reaction-diffusion
model for single species
Raed Ali Alkhasawneh
College of Applied Studies and Community Service, Imam Abdulrahman Bin Faisal University, Saudi Arabia
Article Info ABSTRACT
Article history:
Received Mar 13, 2020
Revised Jun 6, 2020
Accepted Jun 16, 2020
In this paper, we derived a delay advection reaction-diffusion equation with
linear advection term from a stage-structured model, then the derived
equation is used under the homogeneous Dirichlet boundary conditions
𝑒 π‘š(0, 𝑑) = 0, 𝑒 π‘š(𝐿, 𝑑) = 0, and the initial condition 𝑒 π‘š(π‘₯, 0) = 𝑒 π‘š
0 (π‘₯)
> 0, π‘₯ ∈ [βˆ’πœ, 0] with 𝑒 π‘š
0
(0) > 0 in order to find the minimum value of
domain 𝐿 that prevents extinction of the species under the effect of advection
reaction-diffusion equation. Finally, for the measurement the time lengths
from birth to the development of the species population, time delays
are integrated.
Keywords:
Competition
Stage structure
Time delay
Advection
Travelling wave solutions Copyright Β© 2020 Institute of Advanced Engineering and Science.
All rights reserved.
Corresponding Author:
Raed Ali Alkhasawneh,
Department of General Courses,
Imam Abdulrahman Bin Faisal University,
P. O. Box 1982, Dammam, 31441, Saudi Arabia.
Email: ralkhasawneh@iau.edu.sa
1. INTRODUCTION
Reaction-diffusion conditions demonstrate are exceptionally critical in numerous individuals’
developments in an environment or media and nearby response energy such as birth, passing and other
response handle, see Britton [1], Lewis [2], Allen [3] and Aiello [4]. Besides, reaction-diffusion frameworks
can be clarified the impacts of the measure, shape and heterogeneity of the spatial environment on
the diligence of species and the structure of communities in environment. Reaction (𝑓(𝑒)) and dispersal
(spread coefficient 𝐷) both contribute to the inquisitively dynamical behavior of the arrangements of
the condition. We are able to describe the over component by the classical FKPP (Fisher’s condition)
reaction-diffusion condition [5, 6]:
πœ•π‘’
πœ•π‘‘
= 𝑓(𝑒) + 𝐷
πœ•2
𝑒
πœ•π‘₯2
(1)
where 𝑓(𝑒) = 𝑒(1 βˆ’ 𝑒).
It can consider the diffusion as an irregular step which begins at a point and takes place in an
arbitrary heading. Fick’s laws can be utilized to solve for the diffusion coefficient 𝐷 and it addresses that
the diffusive flux goes from areas of high concentration to areas of low concentration. The extraordinary
arrangement of the (1) could be an engendering front (moreover called traveling wave arrangement), the two
non-equilibrium homogeneous states are isolated, one of which (𝑒 = 1) is steady and another one (𝑒 = 0)
is unsteady [7-9]. For illustration, Gurney and Nisbet [8] expanded the collection of stage-structure models
which can be portrayed in terms of delay-differential conditions by analyzing models where the processes of
development and improvement within a stage are distinct. This grants the utilize of delay-differential
Int J Elec & Comp Eng ISSN: 2088-8708 
A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh)
6261
condition models in circumstances where both populace numbers and add up to biomass are powerfully
critical. Al-Omari and Gourley [10, 11] proposed a delay differential condition show for a single species with
stage-structure in which the development delay is displayed as a dissemination, to permit for the possibility
that people may take diverse sums of time to develop. Common birth and passing rate capacities are utilized.
They found that the elements of the demonstrate depends to a great extent on the qualitative frame of
the birth work, which depends on the overall number of grown-ups. They also propose an alternative model
which derived by Aiello, Freedman and Wu, see [12], allowing a non-monotone birth function of the kind
seen in other frequently studied population models. Then they present results on positivity and boundedness,
existence and local stability of equilibria, global stability of the extinction state under certain conditions, and
existence of periodic solutions. Gourley et al., [13] presented demonstrate circuitous transmission, through
contact with infections, of avian influenza in transitory and nonmigratory fowls, taking under consideration
age structure. Relocation is modeled through a reaction-advection condition on a closed circle parametrized
by bend length (the migration flyway) that begins and closes at the area where fowls breed in summer.
Their modeling keeps the winged creatures together as a run, the position of which is certainly decided and
known for all future time. Births happen when the run passes the breeding area and are modeled utilizing
thoughts of rash differential conditions. Sarfaraz [14] classified the parameter spaces for reaction-diffusion
frameworks of two chemical species on stationary spaces in terms of the sorts and solidness of the uniform
consistent state and irritated the impact on the flow of the re-action-diffusion framework.
Chunwei [15] defined a postponed reaction-diffusion demonstrate that depicts competition between
two species in a stream. He isolates each species into two champers, people in-habiting on the seabed and
people floating within the stream. Time delays are consolidated to degree the time lengths from birth
to development of the benthic populaces. It too ponders the elements of the non-spatial show, decide
the presence and worldwide solidness of the equilibrium, and give conditions beneath which arrangements
merge to the balance. At that point he appears that the presence of traveling wave arrangements can be set up
through compact fundamentally operators. At last, he defines two genuine numbers and demonstrate that they
serve as the lower bounds of the speeds of traveling wave arrangements within the framework.
Rattanakul [16] built an advection diffusion-reaction model demonstrate for the angle populace
structured to track the populace densities of both the labeled angle and the tag-free angle, in which the rash
labeling hone is joined, whereas nonstop labeling is accepted to be done on off springs of tractable labeled
angle, or on those within the same swarm as the tractable labeled angle. Utilizing the traveling wave
facilitate, they infer explanatory expression for the arrangements to the show framework. Also, they inferred
the unequivocal expression for the level of labeled angle which increments in an intermittent rash design.
Solidness and stage plane investigations are moreover carried out to determine. While Venkataraman [17]
investigate a model for biological pattern formation during growth development. The pattern formation
phenomenon is described by a reaction-diffusion system on a time-dependent domain.
Boonrangsiman et al. [18] considered an application to fisheries and accepted that there's a single
prey populace and a predator populace that can be isolated by generation capacity into a youthful and
a develop arrange, with a time delay for the youthful to develop move. They demonstrated that the framework
has three nonnegative balance focuses, specifically, an unimportant point with all populaces zero, a predator-
free harmony point, and a coexistence balance point with all three populaces non-zero. It is demonstrated that
the minor harmony point is continuously unsteady, that the predator-free balance point is steady on the off
chance that and as it were in the event that the existence harmony point does not exist, which the existence
point can either be steady for all time delays or ended up unsteady in the event that a Hopf bifurcation occurs
at a basic time delay. While Zhang [19] explored a stage-structured postponed reaction-diffusion show with
advection that depicts competition between two develop species in water stream. Time delays are
consolidated to degree the time lengths from birth to development of the populaces. They appear that there
exists a limited positive number c* that can be characterized as the slowest spreading speed of traveling wave
arrangements interfacing two mono-culture equilibria or interfacing a monoculture with the coexistence
harmony. Moreover, Cangiani et al. [20] illustrated a few modern sorts of wave proliferation and design
arrangement in a classical three species cyclic competition show with spatial dissemination These unused
designs are characterized by a tall normality in space but are diverse from designs already known to exist in
reaction–diffusion models and demonstrated a dependable bound for the blunder of the numerical strategy
which permit us to effectively investigate the dynamical designs in both two and three spatial measurements.
At last, Murray [21-23] examined the condition for the linear stability of 𝑒 = 0 for the taking after response
dissemination demonstrate:
πœ•π‘’
πœ•π‘‘
= 𝑓(𝑒) + 𝐷
πœ•2
𝑒
πœ•π‘₯2
, 0 ≀ π‘₯ ≀ 𝐿, (2)
 ISSN: 2088-8708
Int J Elec & Comp Eng, Vol. 10, No. 6, December 2020 : 6260 - 6267
6262
With the following boundary and initial conditions:
𝑒(π‘₯, 𝑑) = 0, 𝑒(𝐿, 𝑑) = 0, 𝑒(π‘₯, 0) = 𝑒₀(π‘₯),
where 𝑒 denotes the population density of a species, 𝑓(𝑒) is the species dynamics and 𝐷 is the diffusion
coefficient which measure the dispersal efficiency of the relevant species, 𝐿 is the length of the domain.
He considered the different boundary conditions which consider the influence of the region exterior to
the reaction diffusion domain. He showed that if the domain size is not large enough, then (2) cannot
generate spatial patterns and so he found the condition on 𝐿 such that the species will not become extinct if:
𝐿 > πœ‹βˆš
𝐷
𝑓′(0)
In this paper, an age-structured model is used to derive a delay advection reaction-diffusion equation with
linear advection term. In addition, we find the minimum value of 𝐿 that prevents extinction of the species
under the effect of advection term for the reaction-diffusion equation.
2. A DELAY ADVECTION REACTION-DIFFUSION EQUATION WITH LINEAR ADVECTION
TERM
In this section, we derive a delay advection reaction-diffusion equation with linear advection term
from an age-structured model, and then later on, we study and analyze our new model under the effect of
the advection term. Define 𝑒 π‘š(π‘₯, 𝑑) to be the intensity of mature adults, with age of at least Ο„. Then it can be
represented as follows:
𝑒 π‘š(π‘₯, 𝑑) = ∫ 𝑒(π‘₯, 𝑑, π‘Ž)π‘‘π‘Ž
∞
𝜏
, (3)
where 𝑒(π‘₯, 𝑑, π‘Ž) is the intensity of individuals of age π‘Ž at point π‘₯, time 𝑑 and 𝜏 is the length of the juvenile
period. Now, let 𝑒 satisfies the following equation, see Metz [24] and McNair [25]:
πœ•π‘’
πœ•π‘‘
+
πœ•π‘’
πœ•π‘Ž
= π‘ˆ
πœ•π‘’
πœ•π‘₯
+ 𝑑𝑖
πœ•2
𝑒
πœ•π‘₯2
βˆ’ 𝛾𝑒, 0 < π‘Ž < 𝜏, (4)
and that the mature evolve according to:
πœ•π‘’ π‘š
πœ•π‘‘
= π‘ˆ
πœ•π‘’ π‘š
πœ•π‘₯
+ 𝑑 π‘š
πœ•2
𝑒 π‘š
πœ•π‘₯2
+ 𝑒(π‘₯, 𝑑, 𝜏) βˆ’ 𝛽𝑒 π‘š
2
, π‘₯ ∈ (βˆ’βˆž, ∞), 𝑑 > 0 (5)
with
𝑒(π‘₯, 𝑑, 0) = 𝑓(𝑒 π‘š(π‘₯, 𝑑)) (6)
the term 𝑒(π‘₯, 𝑑, 𝜏) in (5) represents adult recruitment, since it is those of age 𝜏 (maturation age) and 𝑓(𝑒 π‘š)
in (6) refers to the rare of the birth, supposed to depend only on the number of matures at the time.
Let 𝑣(π‘₯, π‘Ÿ, π‘Ž) = 𝑒(π‘₯, π‘Ž + π‘Ÿ, π‘Ž), then
πœ•π‘£
πœ•π‘Ž
= [
πœ•π‘’
πœ•π‘‘
(π‘₯, 𝑑, π‘Ž) +
πœ•π‘’
πœ•π‘Ž
(π‘₯, 𝑑, π‘Ž)]
𝑑=π‘Ž+π‘Ÿ
= π‘ˆ
πœ•π‘’
πœ•π‘₯
(π‘₯, π‘Ž + π‘Ÿ, π‘Ž) + 𝑑𝑖
πœ•2
𝑒
πœ•π‘₯2
(π‘₯, π‘Ž + π‘Ÿ, π‘Ž) βˆ’ 𝛾𝑒(π‘₯, π‘Ž + π‘Ÿ, π‘Ž),
which implies
πœ•π‘£
πœ•π‘Ž
= π‘ˆ
πœ•π‘£
πœ•π‘₯
+ 𝑑𝑖
πœ•2
𝑣
πœ•π‘₯2
βˆ’ 𝛾𝑣
πœ•π‘’
πœ•π‘‘
+
πœ•π‘’
πœ•π‘Ž
= π‘ˆ
πœ•π‘’
πœ•π‘₯
+ 𝑑𝑖
πœ•2
𝑒
πœ•π‘₯2
βˆ’ 𝛾𝑒, 0 < π‘Ž < 𝜏, (7)
Int J Elec & Comp Eng ISSN: 2088-8708 
A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh)
6263
Now, we apply the Fourier transform:
𝑣̂(𝑠, π‘Ÿ, π‘Ž) = ∫ 𝑣(π‘₯, π‘Ÿ, π‘Ž) π‘’βˆ’π‘–π‘ π‘₯
𝑑π‘₯
∞
βˆ’βˆž
(8)
To (8) gives:
πœ•π‘£Μ‚
πœ•π‘Ž
= (π‘ˆπ‘–π‘  βˆ’ 𝑑𝑖 𝑠² βˆ’ 𝛾)𝑣̂ (9)
and
𝑣̂(𝑠, π‘Ÿ, 0) = 𝑒̂(𝑠, π‘Ÿ, 0) = β„±{𝑓(𝑒 π‘š(π‘₯, π‘Ÿ))} (10)
The solution of (9) is:
𝑣̂(𝑠, π‘Ÿ, π‘Ž) = β„±{𝑓(𝑒 π‘š(π‘₯, π‘Ÿ))}𝑒(π‘ˆπ‘–π‘ βˆ’π‘‘ 𝑖 π‘ Β²βˆ’π›Ύ)π‘Ž
. (11)
Now,
ℱ⁻¹(𝑒(π‘ˆπ‘–π‘ βˆ’π‘‘ 𝑖 π‘ Β²βˆ’π›Ύ)π‘Ž
) =
1
2πœ‹
𝑒(π‘ˆπ‘–π‘ βˆ’π‘‘ 𝑖 π‘ Β²βˆ’π›Ύ)π‘Ž
𝑒 𝑖𝑠π‘₯
𝑑𝑠
=
π‘’βˆ’π›Ύπ‘Ž
√4πœ‹π‘‘π‘– π‘Ž
π‘’βˆ’(π‘₯+π‘ˆπ‘Ž)Β²/(4𝑑 𝑖 π‘Ž)
.
By using convolution theorem:
𝑣(π‘₯, π‘Ÿ, π‘Ž) = π‘’βˆ’π›Ύπ‘Ž
∫
1
√4πœ‹π‘‘π‘– π‘Ž
∞
βˆ’βˆž
𝑒
βˆ’(π‘ˆπ‘Ž +π‘₯βˆ’π‘¦)2
4𝑑 𝑖 π‘Ž 𝑓(𝑒 π‘š(𝑦, π‘Ÿ))𝑑𝑦,
Therefore
𝑒(π‘₯, 𝑑, 𝜏) = 𝑣(π‘₯, 𝑑 βˆ’ 𝜏, 𝜏)
= π‘’βˆ’π›Ύπœ
∫
1
√4πœ‹π‘‘π‘– 𝜏
∞
βˆ’βˆž
𝑒
βˆ’(π‘ˆπœ +π‘₯βˆ’π‘¦)2
4𝑑 𝑖 𝜏 𝑓(𝑒 π‘š(𝑦, 𝑑 βˆ’ 𝜏))𝑑𝑦. (12)
If 𝑓(𝑒 π‘š) = 𝛼𝑒 π‘š, then the birth rate is symmetric to the number of the mature adults given by 𝛼𝑒 π‘š. Most of
the populations, such presumption is valid only when population numbers are relatively little or when there is
abundance of nourishment. Inserting (12) with 𝑓(𝑒 π‘š) = 𝛼𝑒 π‘š into (4) yields:
πœ•π‘’ π‘š
πœ•π‘‘
= π‘ˆ
πœ•π‘’ π‘š
πœ•π‘₯
+ 𝑑 π‘š
πœ•2
𝑒 π‘š
πœ•π‘₯2
βˆ’ 𝛽𝑒 π‘š
2 +∝ π‘’βˆ’π›Ύπœ ∫
1
√4πœ‹π‘‘π‘– 𝜏
∞
βˆ’βˆž
𝑒
βˆ’(π‘ˆπœ +π‘₯βˆ’π‘¦)2
4𝑑 𝑖 𝜏 𝑒 π‘š(𝑦, 𝑑 βˆ’ 𝜏)𝑑𝑦 (13)
By applying the following substitution in (13):
π‘ˆπœ + π‘₯ βˆ’ 𝑦 = π‘€βˆš4𝑑𝑖 𝜏 ,
we obtain:
π›Όπ‘’βˆ’π›Ύπœ
∫
1
√4πœ‹π‘‘π‘– 𝜏
∞
βˆ’βˆž
𝑒
βˆ’(π‘ˆπœ +π‘₯βˆ’π‘¦)2
4𝑑 𝑖 𝜏 𝑒 π‘š(𝑦, 𝑑 βˆ’ 𝜏)𝑑𝑦
= π›Όπ‘’βˆ’π›Ύπœ
∫
1
√ πœ‹
∞
βˆ’βˆž
π‘’βˆ’π‘€2
𝑒 π‘š(π‘ˆπœ + π‘₯ βˆ’ π‘€βˆš4𝑑𝑖 𝜏, 𝑑 βˆ’ 𝜏)𝑑𝑀.
(14)
This form of the integral allows us to make approximation for small values of the parameters. Let us
approximate (14) for small values of the immature diffusivity 𝑑𝑖:
 ISSN: 2088-8708
Int J Elec & Comp Eng, Vol. 10, No. 6, December 2020 : 6260 - 6267
6264
𝑒 π‘š(π‘₯ + π‘ˆπœ βˆ’ π‘€βˆš4𝑑𝑖 𝜏, 𝑑 βˆ’ 𝜏)
= 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) βˆ’ π‘€βˆš4𝑑𝑖 𝜏
πœ•π‘’ π‘š
πœ•π‘₯
(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) + 2𝑑𝑖 πœπ‘€2
πœ•2
𝑒 π‘š
πœ•π‘₯2
(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏)
Therefore, the integral term in (14) can be written as follows:
π›Όπ‘’βˆ’π›Ύπœ
∫
1
√ πœ‹
∞
βˆ’βˆž
π‘’βˆ’π‘€2
(𝑒 π‘š(π‘₯ + π‘ˆπœ βˆ’ π‘€βˆš4𝑑𝑖 𝜏, 𝑑 βˆ’ 𝜏)) 𝑑𝑀
= π›Όπ‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘ˆπœ + π‘₯, 𝑑 βˆ’ 𝜏) + π›Όπ‘’βˆ’π›Ύπœ
𝑑𝑖 𝜏
πœ•2
𝑒 π‘š
πœ•π‘₯2
(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏)),
(15)
where we have used the following fact:
∫ π‘€Β²π‘’βˆ’π‘€2
𝑑𝑀 =
1
2
√ πœ‹
∞
βˆ’βˆž
.
In the limiting case 𝑑𝑖 β†’ 0 (i.e., immatures not diffusing at all), (13) becomes:
πœ•π‘’ π‘š
πœ•π‘‘
= π‘ˆ
πœ•π‘’ π‘š
πœ•π‘₯
+ 𝑑 π‘š
πœ•2
𝑒 π‘š
πœ•π‘₯2
+ π‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) βˆ’ 𝛽𝑒2
π‘š
(16)
In this case 𝑑𝑖 = 0, the term π›Όπ‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) can be understood ecologically as follows:
Setting 𝑑𝑖 = 0 means the immature are not diffusing. They are, however, still subject to advection. During
the period of immaturity, which is of duration 𝜏, each immature will have moved a distance equal to π‘ˆπœ.
It follows that an individual that becomes mature at location x must have been born at location π‘₯ + π‘ˆπœ (since
the advection is from right to left). The term: π›Όπ‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏), which is the rate at time 𝑑, position
π‘₯, at which individuals become mature, is given simply by the birth rate 𝜏 time units ago at location π‘₯ + π‘ˆπœ,
𝛼𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏), corrected for juvenile mortality which explains the π‘’βˆ’π›Ύπœ
factor. If the species is close to
extinction and the immature are not diffusing, then (16) becomes:
πœ•π‘’ π‘š
πœ•π‘‘
= π‘ˆ
πœ•π‘’ π‘š
πœ•π‘₯
+ 𝑑 π‘š
πœ•2
𝑒 π‘š
πœ•π‘₯2
+ π›Όπ‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏), 0 < π‘₯ < 𝐿, 𝑑 > 0. (17)
3. EFFECT OF ADVECTION TERM ON THE DERIVED DELAY ADVECTION REACTION-
DIFFUSION EQUATION
Starting by studying (17) on homogeneous Dirichlet boundary conditions: 𝑒 π‘š(0, 𝑑) = 0,
𝑒 π‘š(𝐿, 𝑑) = 0, and the initial condition 𝑒 π‘š(π‘₯, 0) = 𝑒 π‘š
0
(π‘₯), π‘₯ ∈ [βˆ’πœ, 0] with 𝑒 π‘š
0
(0) > 0. As a starting point,
we shall also consider the case when the juveniles are not subject to advection, in this case, the term 𝑒 π‘š(π‘₯ +
π‘ˆπœ, 𝑑 βˆ’ 𝜏) of (16) becomes: π›Όπ‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘₯, 𝑑 βˆ’ 𝜏). However, the adults will still be subject to advection.
Seeking trial solutions for (16) of the form 𝑒 π‘š(π‘₯, 𝑑) = 𝑒στ
πœ™(π‘₯), yields:
𝑑 π‘š πœ™β€²β€²(π‘₯) + π‘ˆ πœ™β€²βˆ’π›Ύπœ
π‘’βˆ’πœŽπœ
βˆ’ 𝜎)πœ™(π‘₯) = 0 (18)
The characteristic equation for (18) is
𝑑 π‘š 𝑀² + π‘ˆ 𝑀 + (π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’πœŽπœ
βˆ’ 𝜎) = 0 (19)
and the roots are given by:
𝑀₁, 𝑀₂ =
1
2𝑑 π‘š
(βˆ’π‘ˆ Β± (βˆšπ‘ˆ2 βˆ’ 4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) )
Thus, we have two cases for the roots:
If 𝑀₁ and 𝑀₂ are real, then the solution for πœ™ satisfying the boundary condition πœ™(0) = 0 will be in the form
πœ™(π‘₯) = 𝐴(𝑒 𝑀₁π‘₯
βˆ’ 𝑒 𝑀2 π‘₯
)
Int J Elec & Comp Eng ISSN: 2088-8708 
A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh)
6265
and in order to satisfy πœ™(𝐿) = 0, we will have the zero solution.
If M₁ and Mβ‚‚ are complex, then the solution for πœ™(π‘₯) satisfying the boundary condition πœ™(0) = 0
will be of the form
πœ™(π‘₯) = 𝑒
(
βˆ’π‘ˆπ‘₯
2𝑑 π‘š
)
𝐡 𝑠𝑖𝑛(√
4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) βˆ’ π‘ˆΒ²
2𝑑 π‘š
π‘₯). (20)
In order to be able to satisfy πœ™(𝐿) = 0, we need:
√(4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) βˆ’ π‘ˆΒ²)
2𝑑 π‘š
=
π‘›πœ‹
𝐿
, 𝑛 = 1, 2, 3, ….
That is:
4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’πœŽπœ
βˆ’ 𝜎) βˆ’ π‘ˆ2
=
4𝑑 π‘š
2 𝑛2 πœ‹2
𝐿2 , 𝑛 = 1, 2, 3, …,
which gives the following equation to be solved for 𝜎:
𝜎 = π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’πœŽπœ
βˆ’
π‘ˆΒ²
4𝑑 π‘š
βˆ’
𝑑 π‘š π‘›Β²πœ‹Β²
𝐿²
(21)
The solution 𝑒 π‘š(π‘₯, 𝑑) of the linearized (17) is of the following form
𝑒 π‘š(π‘₯, 𝑑) = 𝑒 πœŽπ‘‘
𝐡𝑛 𝑒
βˆ’π‘ˆπ‘₯
2𝑑 π‘š 𝑠𝑖𝑛 (
𝑛 πœ‹π‘₯
𝐿
) , 𝑛 = 1, 2, 3, …,
where 𝜎 satisfies (20).
Now, we are going to find the solution of (17) satisfying the initial data by summing over 𝑛
to obtain:
𝑒 π‘š(π‘₯, 𝑑) = βˆ‘ 𝐡𝑛 𝑒 𝜎 𝑛 𝑑
∞
𝑛=1
𝑒
βˆ’π‘ˆπ‘₯
2𝑑 π‘š 𝑠𝑖𝑛 (
𝑛 πœ‹π‘₯
𝐿
), (22)
where 𝜎 𝑛 satisfies (21) and using the initial condition and Fourier coefficients, then the value of 𝐡𝑛 is
given by:
𝐡{𝑛} =
2
𝐿
∫ 𝑒 π‘š
0 (π‘₯) 𝑒
π‘ˆπ‘₯
2𝑑 π‘š 𝑠𝑖𝑛 (
𝑛 πœ‹π‘₯
𝐿
) 𝑑π‘₯
𝐿
0
, 𝑛 = 1,2,3, … (23)
Substituting this value in (22) yields the final solution of the linearized problem:
𝑒 π‘š(π‘₯, 𝑑) = βˆ‘ (
2
𝐿
∫ 𝑒 π‘š
0 (π‘₯) 𝑒
π‘ˆπ‘₯
2𝑑 π‘š 𝑠𝑖𝑛 (
𝑛 πœ‹π‘₯
𝐿
) 𝑑π‘₯ Γ—
𝐿
0
∞
𝑛=1 𝑒 𝜎 𝑛 𝑑
𝑒
βˆ’π‘ˆπ‘₯
2𝑑 π‘š 𝑠𝑖𝑛 (
𝑛 πœ‹π‘₯
𝐿
) (24)
The population will decay to zero if π‘…π‘’πœŽ 𝑛 < 0, for all 𝑛 = 1,2,3, …, where 𝜎 𝑛 is any root of (21).
In other words, 𝜎 𝑛satisfies:
𝜎 𝑛 = π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’πœŽ 𝑛 𝑑
βˆ’
𝑑 π‘š 𝑛2
πœ‹2
𝐿2
βˆ’
π‘ˆΒ²
4𝑑 π‘š
In order to detect a possible change of linear stability of the zero solution of (17) we set 𝜎 = π‘–πœ”,
which gives:
π‘ˆ2
4𝑑 π‘š
+
𝑑 π‘š 𝑛2 πœ‹2
𝐿2 + 𝑖𝑀 = π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’π‘–π‘€π‘‘
,
 ISSN: 2088-8708
Int J Elec & Comp Eng, Vol. 10, No. 6, December 2020 : 6260 - 6267
6266
and the complex conjugate of above equation is:
π‘ˆ2
4𝑑 π‘š
+
𝑑 π‘š 𝑛2 πœ‹2
𝐿2 βˆ’ 𝑖𝑀 = π›Όπ‘’βˆ’π›Ύπœ
𝑒 𝑖𝑀𝑑
,
Thus, we are eliminating π‘’βˆ’π‘–πœ”πœ
from the last two equations, yields:
πœ”Β² = π›ΌΒ²π‘’βˆ’2π›Ύπœ
βˆ’ [
π‘ˆ2
4𝑑 π‘š
+
𝑑 π‘š 𝑛2
πœ‹2
𝐿2
]Β².
To find the conditions on the parameters which ensure that: πœ”Β² < 0, since this means that roots of
(21) of the form 𝜎 = π‘–πœ”, with πœ” real, cannot exist. Note that: If πœ”Β² < 0 when n=1 then automatically
πœ”Β² < 0 for n=2, 3, 4, …, therefore it is sufficient to set 𝑛 = 1. When 𝑛 = 1, πœ”Β² < 0 and if the following
inequality holds:
π›Όπ‘’βˆ’π›Ύπœ
<
π‘ˆ2
4𝑑 π‘š
+
𝑑 π‘š πœ‹2
𝐿2
then 𝑒 π‘š(π‘₯, 𝑑) β†’ 0 as 𝑑 β†’ 0 , thus the population will become extinct. So, the species will become extinct in
this case if:
π‘ˆ (advection speed) is large
𝑑 π‘š (mature diffusion) is either very small or very large
𝐿 (length of domain) is small
𝛼 (birth rate) is too small
𝛾 (juvenile mortality) is large
𝜏 (length of juvenile period) is large.
Also, if the following inequality holds:
π›Όπ‘’βˆ’π›Ύπœ
>
π‘ˆ2
4𝑑 π‘š
+
𝑑 π‘š πœ‹2
𝐿2 ,
then 𝜎 𝑛 = π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’πœŽ 𝑛 𝑑
βˆ’
𝑑 π‘š 𝑛2 πœ‹2
𝐿2 βˆ’
π‘ˆΒ²
4𝑑 π‘š
with 𝑛 = 1 has a real positive root 𝜎. This can be shown by drawing
the graphs of
π‘ˆ2
4𝑑 π‘š
+
𝑑 π‘š πœ‹2
𝐿2 + 𝜎 and π›Όπ‘’βˆ’π›Ύπœ
π‘’βˆ’πœŽπ‘‘
against 𝜎, where we can see that the two functions have a real
positive root Οƒ which means that the zero solution of (17) is unstable. So, we can conclude that
the population will survive in this case if
π‘ˆ (advection speed) is too small.
𝐿 (domain) is too large.
𝛼 (birth rate) is too large.
4. DISCUSSIONS AND CONCLUSION
We have derived a delay advection reaction-diffusion equation with linear advection term from an
age-structured model which is given by:
πœ•π‘’ π‘š
πœ•π‘‘
= π‘ˆ
πœ•π‘’ π‘š
πœ•π‘₯
+ 𝑑 π‘š
πœ•2
𝑒 π‘š
πœ•π‘₯2
+ π‘’βˆ’π›Ύπœ
𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) βˆ’ 𝛽𝑒2
π‘š
(25)
and our derived equation as in (24) is studied when the species is close to extinction and the immature are not
diffusing on homogeneous Dirichlet boundary conditions 𝑒 π‘š(0, 𝑑) = 0, 𝑒 π‘š(𝐿, 𝑑) = 0 and the initial condition
𝑒 π‘š(π‘₯, 0) = 𝑒 π‘š
0
β‰₯ 0 for π‘₯ ∈ [βˆ’πœ, 0]. Also, we considered the case when juveniles are not subject to
advection, so the term 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) in (24) becomes 𝑒 π‘š(π‘₯, 𝑑 βˆ’ 𝜏). However, the adults will still be
subject to advection. The conditions on the parameters for the final solution 𝑒 π‘š(π‘₯, 𝑑) for the linearized (25)
that prevents extinction of the species under the defect of advection for the reaction-diffusion are founds,
where the conclusion can be summarized as follows:
ο€­ The species will become extinct if the advection speed if large, mature diffusion is either very small or
very large, length of domain is small, juvenile mortality is large and length of juvenile is large.
ο€­ The species will survive if advection speed is small, domain is too large and birth rate is too large.
Int J Elec & Comp Eng ISSN: 2088-8708 
A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh)
6267
REFERENCES
[1] Britton, N., β€œReaction-Diffusion Equations and Their Applications to Biology,” Academic Press, 1986.
[2] Lewis, M., Petrovskii, S., Potts, J., β€œReaction-Diffusion Models: Single Species,” The Mathematics Behind
Biological Invasions, pp. 69-105, 2016.
[3] Allen, L., β€œPersistence and extinction in single-species reaction-diffusion models,” Bulletin of Mathematical
Bioloy, vol. 45, pp. 209-227, 1983.
[4] Aiello, W. G. and Freedman, H. I., β€œA time-delay model of single species growth with stage structure,”
Math. Biosci., vol. 101, pp. 139-153, 1990.
[5] Fisher, R. A., β€œThe wave of advance of advantageous genes,” Annals of Eugenics, vol. 7, pp. 355-369, 1937.
[6] Kolmogorov, A. Petrovskii, I. and Piscounov, N., β€œA study of the diffusion equation with increase in the amount of
substance, and its application to a biological problem,” In V. M. Tikhomirov, editor, Selected Works of A. N.
Kolmogorov I, pages 248C270. Kluwer 1991. Translated by V. M. Volosov from Bull. Moscow Univ., Math.
Mech. 1, pp. 1-25, 1937.
[7] Fisher R. A., β€œThe genetical theory of natural selection,” Oxford University Press, USA, New Ed edition 2000,
variorum edition 1999, 1930.
[8] Gurney, W. S. C. and Nisbet, R. M., β€œFluctuation periodicity, generation separation, and the expression of Larval
competition,” J. Theoretical Popul. Biol., vol. 28, no. 2, pp. 150-180, 1985.
[9] Ablowitz, M. J. and Zeppetella, A., β€œExplicit solutions of Fisher’s equation for a special wave speed,” Bull. Math.
Biol., vol. 41, pp. 835-840, 1979.
[10] Al-Omari, J. F. M. and Gourley S. A., β€œA nonlocal reaction-diffusion model for a single species with stage
structure and distributed maturation delay,” Euro. J. Appl. Math., vol. 16, no. 1, pp. 37-51, 2005.
[11] Al-Omari, J., Gourley, S., β€œDynamics of a stage-structured population model incorporating a state-dependent
maturation delay,” Nonlinear Anal. Real World Appl., vol. 6, no. 1, pp. 13-33, 2005.
[12] Aiello, W. G., Freedman, H. I. and Wu, J., β€œAnalysis of a model representing stage-structured population growth
with state-dependent time delay,” SIAM J. Appl. Math., vol. 52, pp. 855-869, 1992.
[13] Gourley, S. A., Jennings, R. and Liu, R., β€œAn Advection and Age-Structured Approach to Modeling Bird Migration
and Indirect Transmission of Avian Influenza,” SIAM J. Appl. Math., vol. 75, no. 4, pp. 1620-1647, 2005.
[14] Sarfaraz W, Madzvamuse A., β€œClassification of parameter spaces for a reaction diffusion model onstationary
domains,” Chaos Solitons Fractals, vol. 103, pp. 33-51, 2007.
[15] Chunwei W., β€œA stage-structured delayed reaction-diffusion model for A stage-structured delayed reaction-
diffusion model for competition between two species,” Electronic Theses and Dissertations, University of
Louisville, 2013.
[16] Rattanakul, C., Lenbury, Y. and Suksamran, J., β€œAnalysis of advection-diffusion-reaction model for fish population
movement with impulsive tagging: stability and traveling wave solution,” Advances in Difference Equations,
vol. 2019, 2019.
[17] Venkataraman, C., β€œReaction-diffusion system on evolving dmains with applications to the theory of biological
pattern formation,” Ph.D. thesis, University of Sussex, 2011.
[18] Boonrangsiman, S., Bunwon, K. and Elvin, J., β€œA bifurcation path to chaos in a time-delay fisheries predator-prey model
with prey consumption by immature and mature predators,” Math. Computers Simul., vol. 124, pp. 16-29, 2016.
[19] Zhang, L. and Zhao, H. Y., β€œTraveling wave solutions in a stage-structured delayed reaction-diffusion model with
advection,” J. Math. Modelling Anal., vol. 20, no. 2, pp. 168-187, 2015.
[20] Cangiani, A., Georgoulis, E., Morozov, A. and Sutton O., β€œRevealing new dynamical patterns in a reaction-
diffusion model with cyclic competition via a novel computational framework,” Proc.R.Soc.A474, 2018.
[21] Murray, J. D., β€œMathematical Biology,” 2nd ed., Springer, Berlin, 1993.
[22] Murray, J. D., β€œMathematical Biology,” I. An introduction. 3rd ed. Inter, disciplinary Applied Mathematics,
Springer, New York, 2002.
[23] Murray, J. D. , β€œMathematical biology,” Spatial models and biomedical applications. Springer, New York, 2013.
[24] Metz, J. A. J and Diekmann, O., β€œThe Dynamics of Physiologically Structured Populations,” Springer-Verlag,
New York, 1986.
[25] McNair, J., N., and Goulden, C. E., β€œThe dynamics of age-structured populations with a gestation period: Density-
independent growth and egg ration methods for estimating the birth rate,” Theoret. Population Bio., vol. 39,
pp. 1-29, 1991.

Recomendados

Dynamic Habitat Models for Estuary-Dependent Species por
Dynamic Habitat Models for Estuary-Dependent SpeciesDynamic Habitat Models for Estuary-Dependent Species
Dynamic Habitat Models for Estuary-Dependent SpeciesNisqually River Council
169 visualizaΓ§Γ΅esβ€’39 slides
İn saturated groundwater flow por
İn saturated groundwater flowİn saturated groundwater flow
İn saturated groundwater flowsmailKalafat1
93 visualizaΓ§Γ΅esβ€’46 slides
Turbulence generation by planar detonations in heterogeneous mixtures por
Turbulence generation by planar detonations in heterogeneous mixturesTurbulence generation by planar detonations in heterogeneous mixtures
Turbulence generation by planar detonations in heterogeneous mixturesAlberto Cuadra Lara
19 visualizaΓ§Γ΅esβ€’1 slide
Fractional calculus applied in solving instability phenomenon in fluid dynamics por
Fractional calculus applied in solving instability phenomenon in fluid dynamicsFractional calculus applied in solving instability phenomenon in fluid dynamics
Fractional calculus applied in solving instability phenomenon in fluid dynamicsIAEME Publication
244 visualizaΓ§Γ΅esβ€’11 slides
Paine 1980 food webs por
Paine 1980   food websPaine 1980   food webs
Paine 1980 food websEdinaldo Nelson dos Santos Silva
1.1K visualizaΓ§Γ΅esβ€’21 slides
D0562022 por
D0562022D0562022
D0562022IOSR Journals
291 visualizaΓ§Γ΅esβ€’3 slides

Mais conteΓΊdo relacionado

Similar a A stage-structured delayed advection reaction-diffusion model for single species

The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func... por
The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func...The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func...
The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func...mathsjournal
26 visualizaΓ§Γ΅esβ€’21 slides
document por
documentdocument
documentAdam Zienkiewicz
139 visualizaΓ§Γ΅esβ€’17 slides
As pi re2015_abstracts por
As pi re2015_abstractsAs pi re2015_abstracts
As pi re2015_abstractsJoseph Park
256 visualizaΓ§Γ΅esβ€’8 slides
Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen... por
Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen...Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen...
Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen...IJERA Editor
303 visualizaΓ§Γ΅esβ€’5 slides
The Probability distribution of a Simple Stochastic Infection and Recovery Pr... por
The Probability distribution of a Simple Stochastic Infection and Recovery Pr...The Probability distribution of a Simple Stochastic Infection and Recovery Pr...
The Probability distribution of a Simple Stochastic Infection and Recovery Pr...IOSRJM
65 visualizaΓ§Γ΅esβ€’10 slides
Form, function, and evolution of living organisms por
Form, function, and evolution of living organismsForm, function, and evolution of living organisms
Form, function, and evolution of living organismsJosΓ© Luis Moreno Garvayo
848 visualizaΓ§Γ΅esβ€’6 slides

Similar a A stage-structured delayed advection reaction-diffusion model for single species (20)

The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func... por mathsjournal
The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func...The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func...
The Impact of Allee Effect on a Predator-Prey Model with Holling Type II Func...
mathsjournalβ€’26 visualizaΓ§Γ΅es
document por Adam Zienkiewicz
documentdocument
document
Adam Zienkiewiczβ€’139 visualizaΓ§Γ΅es
As pi re2015_abstracts por Joseph Park
As pi re2015_abstractsAs pi re2015_abstracts
As pi re2015_abstracts
Joseph Parkβ€’256 visualizaΓ§Γ΅es
Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen... por IJERA Editor
Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen...Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen...
Stochastic Model to Find the Gallbladder Motility in Acromegaly Using Exponen...
IJERA Editorβ€’303 visualizaΓ§Γ΅es
The Probability distribution of a Simple Stochastic Infection and Recovery Pr... por IOSRJM
The Probability distribution of a Simple Stochastic Infection and Recovery Pr...The Probability distribution of a Simple Stochastic Infection and Recovery Pr...
The Probability distribution of a Simple Stochastic Infection and Recovery Pr...
IOSRJMβ€’65 visualizaΓ§Γ΅es
Form, function, and evolution of living organisms por JosΓ© Luis Moreno Garvayo
Form, function, and evolution of living organismsForm, function, and evolution of living organisms
Form, function, and evolution of living organisms
JosΓ© Luis Moreno Garvayoβ€’848 visualizaΓ§Γ΅es
Study of reflection and transmission of axially symmetric body waves incident... por NandaKumar118171
Study of reflection and transmission of axially symmetric body waves incident...Study of reflection and transmission of axially symmetric body waves incident...
Study of reflection and transmission of axially symmetric body waves incident...
NandaKumar118171β€’6 visualizaΓ§Γ΅es
Transition for regular to mach reflection hysteresis por eSAT Publishing House
Transition for regular to mach reflection hysteresisTransition for regular to mach reflection hysteresis
Transition for regular to mach reflection hysteresis
eSAT Publishing Houseβ€’298 visualizaΓ§Γ΅es
Annals of Limnology and Oceanography por peertechzpublication
Annals of Limnology and OceanographyAnnals of Limnology and Oceanography
Annals of Limnology and Oceanography
peertechzpublicationβ€’30 visualizaΓ§Γ΅es
A Analysis On Probabilistic Risk por Ann Garcia
A Analysis On Probabilistic RiskA Analysis On Probabilistic Risk
A Analysis On Probabilistic Risk
Ann Garciaβ€’2 visualizaΓ§Γ΅es
System of quasilinear equations of reaction diffusion por eSAT Publishing House
System of quasilinear equations of reaction diffusionSystem of quasilinear equations of reaction diffusion
System of quasilinear equations of reaction diffusion
eSAT Publishing Houseβ€’262 visualizaΓ§Γ΅es
Life Expectancy Estimate with Bivariate Weibull Distribution using Archimedea... por CSCJournals
Life Expectancy Estimate with Bivariate Weibull Distribution using Archimedea...Life Expectancy Estimate with Bivariate Weibull Distribution using Archimedea...
Life Expectancy Estimate with Bivariate Weibull Distribution using Archimedea...
CSCJournalsβ€’177 visualizaΓ§Γ΅es
Physical Characterization of a Method for Production of High Stability Suspen... por Editor IJCATR
Physical Characterization of a Method for Production of High Stability Suspen...Physical Characterization of a Method for Production of High Stability Suspen...
Physical Characterization of a Method for Production of High Stability Suspen...
Editor IJCATRβ€’252 visualizaΓ§Γ΅es
Model And Assumptions Of The Royle Nichols Model por Tiffany Rose
Model And Assumptions Of The Royle Nichols ModelModel And Assumptions Of The Royle Nichols Model
Model And Assumptions Of The Royle Nichols Model
Tiffany Roseβ€’3 visualizaΓ§Γ΅es
Emergent global patterns_of_ecosystem_structure_and_function_form_a_mechanist... por Dr Lendy Spires
Emergent global patterns_of_ecosystem_structure_and_function_form_a_mechanist...Emergent global patterns_of_ecosystem_structure_and_function_form_a_mechanist...
Emergent global patterns_of_ecosystem_structure_and_function_form_a_mechanist...
Dr Lendy Spiresβ€’146 visualizaΓ§Γ΅es
Reservoirs & Graphs por Riccardo Rigon
Reservoirs & GraphsReservoirs & Graphs
Reservoirs & Graphs
Riccardo Rigonβ€’2.3K visualizaΓ§Γ΅es
FRACTIONAL CALCULUS APPLIED IN SOLVING INSTABILITY PHENOMENON IN FLUID DYNAMICS por IAEME Publication
FRACTIONAL CALCULUS APPLIED IN SOLVING INSTABILITY PHENOMENON IN FLUID DYNAMICSFRACTIONAL CALCULUS APPLIED IN SOLVING INSTABILITY PHENOMENON IN FLUID DYNAMICS
FRACTIONAL CALCULUS APPLIED IN SOLVING INSTABILITY PHENOMENON IN FLUID DYNAMICS
IAEME Publicationβ€’422 visualizaΓ§Γ΅es
bioexpo_poster por Chiyu Jiang
bioexpo_posterbioexpo_poster
bioexpo_poster
Chiyu Jiangβ€’85 visualizaΓ§Γ΅es
Nonlinear Darcy flow dynamics during ganglia stranding and mobilization in he... por Anastasia Dollari
Nonlinear Darcy flow dynamics during ganglia stranding and mobilization in he...Nonlinear Darcy flow dynamics during ganglia stranding and mobilization in he...
Nonlinear Darcy flow dynamics during ganglia stranding and mobilization in he...
Anastasia Dollariβ€’80 visualizaΓ§Γ΅es
Reaction Paper On Gels por Aurora Cuellar
Reaction Paper On GelsReaction Paper On Gels
Reaction Paper On Gels
Aurora Cuellarβ€’2 visualizaΓ§Γ΅es

Mais de IJECEIAES

Predictive fertilization models for potato crops using machine learning techn... por
Predictive fertilization models for potato crops using machine learning techn...Predictive fertilization models for potato crops using machine learning techn...
Predictive fertilization models for potato crops using machine learning techn...IJECEIAES
3 visualizaΓ§Γ΅esβ€’9 slides
Computer-aided automated detection of kidney disease using supervised learnin... por
Computer-aided automated detection of kidney disease using supervised learnin...Computer-aided automated detection of kidney disease using supervised learnin...
Computer-aided automated detection of kidney disease using supervised learnin...IJECEIAES
2 visualizaΓ§Γ΅esβ€’10 slides
Reliable and efficient webserver management for task scheduling in edge-cloud... por
Reliable and efficient webserver management for task scheduling in edge-cloud...Reliable and efficient webserver management for task scheduling in edge-cloud...
Reliable and efficient webserver management for task scheduling in edge-cloud...IJECEIAES
2 visualizaΓ§Γ΅esβ€’10 slides
Reinforcement learning-based security schema mitigating manin-the-middle atta... por
Reinforcement learning-based security schema mitigating manin-the-middle atta...Reinforcement learning-based security schema mitigating manin-the-middle atta...
Reinforcement learning-based security schema mitigating manin-the-middle atta...IJECEIAES
2 visualizaΓ§Γ΅esβ€’14 slides
Resource-efficient workload task scheduling for cloud-assisted internet of th... por
Resource-efficient workload task scheduling for cloud-assisted internet of th...Resource-efficient workload task scheduling for cloud-assisted internet of th...
Resource-efficient workload task scheduling for cloud-assisted internet of th...IJECEIAES
2 visualizaΓ§Γ΅esβ€’10 slides
Breast cancer classification with histopathological image based on machine le... por
Breast cancer classification with histopathological image based on machine le...Breast cancer classification with histopathological image based on machine le...
Breast cancer classification with histopathological image based on machine le...IJECEIAES
4 visualizaΓ§Γ΅esβ€’13 slides

Mais de IJECEIAES(20)

Predictive fertilization models for potato crops using machine learning techn... por IJECEIAES
Predictive fertilization models for potato crops using machine learning techn...Predictive fertilization models for potato crops using machine learning techn...
Predictive fertilization models for potato crops using machine learning techn...
IJECEIAESβ€’3 visualizaΓ§Γ΅es
Computer-aided automated detection of kidney disease using supervised learnin... por IJECEIAES
Computer-aided automated detection of kidney disease using supervised learnin...Computer-aided automated detection of kidney disease using supervised learnin...
Computer-aided automated detection of kidney disease using supervised learnin...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Reliable and efficient webserver management for task scheduling in edge-cloud... por IJECEIAES
Reliable and efficient webserver management for task scheduling in edge-cloud...Reliable and efficient webserver management for task scheduling in edge-cloud...
Reliable and efficient webserver management for task scheduling in edge-cloud...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Reinforcement learning-based security schema mitigating manin-the-middle atta... por IJECEIAES
Reinforcement learning-based security schema mitigating manin-the-middle atta...Reinforcement learning-based security schema mitigating manin-the-middle atta...
Reinforcement learning-based security schema mitigating manin-the-middle atta...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Resource-efficient workload task scheduling for cloud-assisted internet of th... por IJECEIAES
Resource-efficient workload task scheduling for cloud-assisted internet of th...Resource-efficient workload task scheduling for cloud-assisted internet of th...
Resource-efficient workload task scheduling for cloud-assisted internet of th...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Breast cancer classification with histopathological image based on machine le... por IJECEIAES
Breast cancer classification with histopathological image based on machine le...Breast cancer classification with histopathological image based on machine le...
Breast cancer classification with histopathological image based on machine le...
IJECEIAESβ€’4 visualizaΓ§Γ΅es
Early detection of dysphoria using electroencephalogram affective modelling por IJECEIAES
Early detection of dysphoria using electroencephalogram affective modelling Early detection of dysphoria using electroencephalogram affective modelling
Early detection of dysphoria using electroencephalogram affective modelling
IJECEIAESβ€’4 visualizaΓ§Γ΅es
A novel defect detection method for software requirements inspections por IJECEIAES
A novel defect detection method for software requirements inspections A novel defect detection method for software requirements inspections
A novel defect detection method for software requirements inspections
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Convolutional auto-encoded extreme learning machine for incremental learning ... por IJECEIAES
Convolutional auto-encoded extreme learning machine for incremental learning ...Convolutional auto-encoded extreme learning machine for incremental learning ...
Convolutional auto-encoded extreme learning machine for incremental learning ...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Brain cone beam computed tomography image analysis using ResNet50 for collate... por IJECEIAES
Brain cone beam computed tomography image analysis using ResNet50 for collate...Brain cone beam computed tomography image analysis using ResNet50 for collate...
Brain cone beam computed tomography image analysis using ResNet50 for collate...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Channel and spatial attention mechanism for fashion image captioning por IJECEIAES
Channel and spatial attention mechanism for fashion image captioning Channel and spatial attention mechanism for fashion image captioning
Channel and spatial attention mechanism for fashion image captioning
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Regional feature learning using attribute structural analysis in bipartite at... por IJECEIAES
Regional feature learning using attribute structural analysis in bipartite at...Regional feature learning using attribute structural analysis in bipartite at...
Regional feature learning using attribute structural analysis in bipartite at...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Adaptive traffic lights based on traffic flow prediction using machine learni... por IJECEIAES
Adaptive traffic lights based on traffic flow prediction using machine learni...Adaptive traffic lights based on traffic flow prediction using machine learni...
Adaptive traffic lights based on traffic flow prediction using machine learni...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Content-based product image retrieval using squared-hinge loss trained convol... por IJECEIAES
Content-based product image retrieval using squared-hinge loss trained convol...Content-based product image retrieval using squared-hinge loss trained convol...
Content-based product image retrieval using squared-hinge loss trained convol...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
A deep convolutional structure-based approach for accurate recognition of ski... por IJECEIAES
A deep convolutional structure-based approach for accurate recognition of ski...A deep convolutional structure-based approach for accurate recognition of ski...
A deep convolutional structure-based approach for accurate recognition of ski...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
IPv6 flood attack detection based on epsilon greedy optimized Q learning in s... por IJECEIAES
IPv6 flood attack detection based on epsilon greedy optimized Q learning in s...IPv6 flood attack detection based on epsilon greedy optimized Q learning in s...
IPv6 flood attack detection based on epsilon greedy optimized Q learning in s...
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Analysis and prediction of seed quality using machine learning por IJECEIAES
Analysis and prediction of seed quality using machine learning Analysis and prediction of seed quality using machine learning
Analysis and prediction of seed quality using machine learning
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Explainable extreme boosting model for breast cancer diagnosis por IJECEIAES
Explainable extreme boosting model for breast cancer diagnosis Explainable extreme boosting model for breast cancer diagnosis
Explainable extreme boosting model for breast cancer diagnosis
IJECEIAESβ€’2 visualizaΓ§Γ΅es
Predicting active compounds for lung cancer based on quantitative structure-a... por IJECEIAES
Predicting active compounds for lung cancer based on quantitative structure-a...Predicting active compounds for lung cancer based on quantitative structure-a...
Predicting active compounds for lung cancer based on quantitative structure-a...
IJECEIAESβ€’4 visualizaΓ§Γ΅es
U-Net transfer learning backbones for lesions segmentation in breast ultrasou... por IJECEIAES
U-Net transfer learning backbones for lesions segmentation in breast ultrasou...U-Net transfer learning backbones for lesions segmentation in breast ultrasou...
U-Net transfer learning backbones for lesions segmentation in breast ultrasou...
IJECEIAESβ€’2 visualizaΓ§Γ΅es

Último

DESIGN OF SPRINGS-UNIT4.pptx por
DESIGN OF SPRINGS-UNIT4.pptxDESIGN OF SPRINGS-UNIT4.pptx
DESIGN OF SPRINGS-UNIT4.pptxgopinathcreddy
21 visualizaΓ§Γ΅esβ€’47 slides
GPS Survery Presentation/ Slides por
GPS Survery Presentation/ SlidesGPS Survery Presentation/ Slides
GPS Survery Presentation/ SlidesOmarFarukEmon1
7 visualizaΓ§Γ΅esβ€’13 slides
unit 1.pptx por
unit 1.pptxunit 1.pptx
unit 1.pptxrrbornarecm
5 visualizaΓ§Γ΅esβ€’53 slides
Renewal Projects in Seismic Construction por
Renewal Projects in Seismic ConstructionRenewal Projects in Seismic Construction
Renewal Projects in Seismic ConstructionEngineering & Seismic Construction
6 visualizaΓ§Γ΅esβ€’8 slides
sam_software_eng_cv.pdf por
sam_software_eng_cv.pdfsam_software_eng_cv.pdf
sam_software_eng_cv.pdfsammyigbinovia
11 visualizaΓ§Γ΅esβ€’5 slides
BCIC - Manufacturing Conclave - Technology-Driven Manufacturing for Growth por
BCIC - Manufacturing Conclave -  Technology-Driven Manufacturing for GrowthBCIC - Manufacturing Conclave -  Technology-Driven Manufacturing for Growth
BCIC - Manufacturing Conclave - Technology-Driven Manufacturing for GrowthInnomantra
20 visualizaΓ§Γ΅esβ€’4 slides

Último(20)

DESIGN OF SPRINGS-UNIT4.pptx por gopinathcreddy
DESIGN OF SPRINGS-UNIT4.pptxDESIGN OF SPRINGS-UNIT4.pptx
DESIGN OF SPRINGS-UNIT4.pptx
gopinathcreddyβ€’21 visualizaΓ§Γ΅es
GPS Survery Presentation/ Slides por OmarFarukEmon1
GPS Survery Presentation/ SlidesGPS Survery Presentation/ Slides
GPS Survery Presentation/ Slides
OmarFarukEmon1β€’7 visualizaΓ§Γ΅es
unit 1.pptx por rrbornarecm
unit 1.pptxunit 1.pptx
unit 1.pptx
rrbornarecmβ€’5 visualizaΓ§Γ΅es
sam_software_eng_cv.pdf por sammyigbinovia
sam_software_eng_cv.pdfsam_software_eng_cv.pdf
sam_software_eng_cv.pdf
sammyigbinoviaβ€’11 visualizaΓ§Γ΅es
BCIC - Manufacturing Conclave - Technology-Driven Manufacturing for Growth por Innomantra
BCIC - Manufacturing Conclave -  Technology-Driven Manufacturing for GrowthBCIC - Manufacturing Conclave -  Technology-Driven Manufacturing for Growth
BCIC - Manufacturing Conclave - Technology-Driven Manufacturing for Growth
Innomantra β€’20 visualizaΓ§Γ΅es
AWS A5.18 A5.18M-2021.pdf por ThinhNguyen455948
AWS A5.18 A5.18M-2021.pdfAWS A5.18 A5.18M-2021.pdf
AWS A5.18 A5.18M-2021.pdf
ThinhNguyen455948β€’8 visualizaΓ§Γ΅es
CPM Schedule Float.pptx por Mathew Joseph
CPM Schedule Float.pptxCPM Schedule Float.pptx
CPM Schedule Float.pptx
Mathew Josephβ€’6 visualizaΓ§Γ΅es
Web Dev Session 1.pptx por VedVekhande
Web Dev Session 1.pptxWeb Dev Session 1.pptx
Web Dev Session 1.pptx
VedVekhandeβ€’20 visualizaΓ§Γ΅es
REACTJS.pdf por ArthyR3
REACTJS.pdfREACTJS.pdf
REACTJS.pdf
ArthyR3β€’37 visualizaΓ§Γ΅es
Integrating Sustainable Development Goals (SDGs) in School Education por SheetalTank1
Integrating Sustainable Development Goals (SDGs) in School EducationIntegrating Sustainable Development Goals (SDGs) in School Education
Integrating Sustainable Development Goals (SDGs) in School Education
SheetalTank1β€’11 visualizaΓ§Γ΅es
Plant Design Report-Oil Refinery.pdf por Safeen Yaseen Ja'far
Plant Design Report-Oil Refinery.pdfPlant Design Report-Oil Refinery.pdf
Plant Design Report-Oil Refinery.pdf
Safeen Yaseen Ja'farβ€’9 visualizaΓ§Γ΅es
2023Dec ASU Wang NETR Group Research Focus and Facility Overview.pptx por lwang78
2023Dec ASU Wang NETR Group Research Focus and Facility Overview.pptx2023Dec ASU Wang NETR Group Research Focus and Facility Overview.pptx
2023Dec ASU Wang NETR Group Research Focus and Facility Overview.pptx
lwang78β€’188 visualizaΓ§Γ΅es
Unlocking Research Visibility.pdf por KhatirNaima
Unlocking Research Visibility.pdfUnlocking Research Visibility.pdf
Unlocking Research Visibility.pdf
KhatirNaimaβ€’11 visualizaΓ§Γ΅es
CrΓ©ativitΓ© dans le design mΓ©canique Γ  l’aide de l’optimisation topologique por LIEGE CREATIVE
CrΓ©ativitΓ© dans le design mΓ©canique Γ  l’aide de l’optimisation topologiqueCrΓ©ativitΓ© dans le design mΓ©canique Γ  l’aide de l’optimisation topologique
CrΓ©ativitΓ© dans le design mΓ©canique Γ  l’aide de l’optimisation topologique
LIEGE CREATIVEβ€’8 visualizaΓ§Γ΅es
GDSC Mikroskil Members Onboarding 2023.pdf por gdscmikroskil
GDSC Mikroskil Members Onboarding 2023.pdfGDSC Mikroskil Members Onboarding 2023.pdf
GDSC Mikroskil Members Onboarding 2023.pdf
gdscmikroskilβ€’68 visualizaΓ§Γ΅es
MongoDB.pdf por ArthyR3
MongoDB.pdfMongoDB.pdf
MongoDB.pdf
ArthyR3β€’51 visualizaΓ§Γ΅es
Ansari: Practical experiences with an LLM-based Islamic Assistant por M Waleed Kadous
Ansari: Practical experiences with an LLM-based Islamic AssistantAnsari: Practical experiences with an LLM-based Islamic Assistant
Ansari: Practical experiences with an LLM-based Islamic Assistant
M Waleed Kadousβ€’11 visualizaΓ§Γ΅es
Design of Structures and Foundations for Vibrating Machines, Arya-ONeill-Pinc... por csegroupvn
Design of Structures and Foundations for Vibrating Machines, Arya-ONeill-Pinc...Design of Structures and Foundations for Vibrating Machines, Arya-ONeill-Pinc...
Design of Structures and Foundations for Vibrating Machines, Arya-ONeill-Pinc...
csegroupvnβ€’13 visualizaΓ§Γ΅es

A stage-structured delayed advection reaction-diffusion model for single species

  • 1. International Journal of Electrical and Computer Engineering (IJECE) Vol. 10, No. 6, December 2020, pp. 6260~6267 ISSN: 2088-8708, DOI: 10.11591/ijece.v10i6.pp6260-6267  6260 Journal homepage: http://ijece.iaescore.com/index.php/IJECE A stage-structured delayed advection reaction-diffusion model for single species Raed Ali Alkhasawneh College of Applied Studies and Community Service, Imam Abdulrahman Bin Faisal University, Saudi Arabia Article Info ABSTRACT Article history: Received Mar 13, 2020 Revised Jun 6, 2020 Accepted Jun 16, 2020 In this paper, we derived a delay advection reaction-diffusion equation with linear advection term from a stage-structured model, then the derived equation is used under the homogeneous Dirichlet boundary conditions 𝑒 π‘š(0, 𝑑) = 0, 𝑒 π‘š(𝐿, 𝑑) = 0, and the initial condition 𝑒 π‘š(π‘₯, 0) = 𝑒 π‘š 0 (π‘₯) > 0, π‘₯ ∈ [βˆ’πœ, 0] with 𝑒 π‘š 0 (0) > 0 in order to find the minimum value of domain 𝐿 that prevents extinction of the species under the effect of advection reaction-diffusion equation. Finally, for the measurement the time lengths from birth to the development of the species population, time delays are integrated. Keywords: Competition Stage structure Time delay Advection Travelling wave solutions Copyright Β© 2020 Institute of Advanced Engineering and Science. All rights reserved. Corresponding Author: Raed Ali Alkhasawneh, Department of General Courses, Imam Abdulrahman Bin Faisal University, P. O. Box 1982, Dammam, 31441, Saudi Arabia. Email: ralkhasawneh@iau.edu.sa 1. INTRODUCTION Reaction-diffusion conditions demonstrate are exceptionally critical in numerous individuals’ developments in an environment or media and nearby response energy such as birth, passing and other response handle, see Britton [1], Lewis [2], Allen [3] and Aiello [4]. Besides, reaction-diffusion frameworks can be clarified the impacts of the measure, shape and heterogeneity of the spatial environment on the diligence of species and the structure of communities in environment. Reaction (𝑓(𝑒)) and dispersal (spread coefficient 𝐷) both contribute to the inquisitively dynamical behavior of the arrangements of the condition. We are able to describe the over component by the classical FKPP (Fisher’s condition) reaction-diffusion condition [5, 6]: πœ•π‘’ πœ•π‘‘ = 𝑓(𝑒) + 𝐷 πœ•2 𝑒 πœ•π‘₯2 (1) where 𝑓(𝑒) = 𝑒(1 βˆ’ 𝑒). It can consider the diffusion as an irregular step which begins at a point and takes place in an arbitrary heading. Fick’s laws can be utilized to solve for the diffusion coefficient 𝐷 and it addresses that the diffusive flux goes from areas of high concentration to areas of low concentration. The extraordinary arrangement of the (1) could be an engendering front (moreover called traveling wave arrangement), the two non-equilibrium homogeneous states are isolated, one of which (𝑒 = 1) is steady and another one (𝑒 = 0) is unsteady [7-9]. For illustration, Gurney and Nisbet [8] expanded the collection of stage-structure models which can be portrayed in terms of delay-differential conditions by analyzing models where the processes of development and improvement within a stage are distinct. This grants the utilize of delay-differential
  • 2. Int J Elec & Comp Eng ISSN: 2088-8708  A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh) 6261 condition models in circumstances where both populace numbers and add up to biomass are powerfully critical. Al-Omari and Gourley [10, 11] proposed a delay differential condition show for a single species with stage-structure in which the development delay is displayed as a dissemination, to permit for the possibility that people may take diverse sums of time to develop. Common birth and passing rate capacities are utilized. They found that the elements of the demonstrate depends to a great extent on the qualitative frame of the birth work, which depends on the overall number of grown-ups. They also propose an alternative model which derived by Aiello, Freedman and Wu, see [12], allowing a non-monotone birth function of the kind seen in other frequently studied population models. Then they present results on positivity and boundedness, existence and local stability of equilibria, global stability of the extinction state under certain conditions, and existence of periodic solutions. Gourley et al., [13] presented demonstrate circuitous transmission, through contact with infections, of avian influenza in transitory and nonmigratory fowls, taking under consideration age structure. Relocation is modeled through a reaction-advection condition on a closed circle parametrized by bend length (the migration flyway) that begins and closes at the area where fowls breed in summer. Their modeling keeps the winged creatures together as a run, the position of which is certainly decided and known for all future time. Births happen when the run passes the breeding area and are modeled utilizing thoughts of rash differential conditions. Sarfaraz [14] classified the parameter spaces for reaction-diffusion frameworks of two chemical species on stationary spaces in terms of the sorts and solidness of the uniform consistent state and irritated the impact on the flow of the re-action-diffusion framework. Chunwei [15] defined a postponed reaction-diffusion demonstrate that depicts competition between two species in a stream. He isolates each species into two champers, people in-habiting on the seabed and people floating within the stream. Time delays are consolidated to degree the time lengths from birth to development of the benthic populaces. It too ponders the elements of the non-spatial show, decide the presence and worldwide solidness of the equilibrium, and give conditions beneath which arrangements merge to the balance. At that point he appears that the presence of traveling wave arrangements can be set up through compact fundamentally operators. At last, he defines two genuine numbers and demonstrate that they serve as the lower bounds of the speeds of traveling wave arrangements within the framework. Rattanakul [16] built an advection diffusion-reaction model demonstrate for the angle populace structured to track the populace densities of both the labeled angle and the tag-free angle, in which the rash labeling hone is joined, whereas nonstop labeling is accepted to be done on off springs of tractable labeled angle, or on those within the same swarm as the tractable labeled angle. Utilizing the traveling wave facilitate, they infer explanatory expression for the arrangements to the show framework. Also, they inferred the unequivocal expression for the level of labeled angle which increments in an intermittent rash design. Solidness and stage plane investigations are moreover carried out to determine. While Venkataraman [17] investigate a model for biological pattern formation during growth development. The pattern formation phenomenon is described by a reaction-diffusion system on a time-dependent domain. Boonrangsiman et al. [18] considered an application to fisheries and accepted that there's a single prey populace and a predator populace that can be isolated by generation capacity into a youthful and a develop arrange, with a time delay for the youthful to develop move. They demonstrated that the framework has three nonnegative balance focuses, specifically, an unimportant point with all populaces zero, a predator- free harmony point, and a coexistence balance point with all three populaces non-zero. It is demonstrated that the minor harmony point is continuously unsteady, that the predator-free balance point is steady on the off chance that and as it were in the event that the existence harmony point does not exist, which the existence point can either be steady for all time delays or ended up unsteady in the event that a Hopf bifurcation occurs at a basic time delay. While Zhang [19] explored a stage-structured postponed reaction-diffusion show with advection that depicts competition between two develop species in water stream. Time delays are consolidated to degree the time lengths from birth to development of the populaces. They appear that there exists a limited positive number c* that can be characterized as the slowest spreading speed of traveling wave arrangements interfacing two mono-culture equilibria or interfacing a monoculture with the coexistence harmony. Moreover, Cangiani et al. [20] illustrated a few modern sorts of wave proliferation and design arrangement in a classical three species cyclic competition show with spatial dissemination These unused designs are characterized by a tall normality in space but are diverse from designs already known to exist in reaction–diffusion models and demonstrated a dependable bound for the blunder of the numerical strategy which permit us to effectively investigate the dynamical designs in both two and three spatial measurements. At last, Murray [21-23] examined the condition for the linear stability of 𝑒 = 0 for the taking after response dissemination demonstrate: πœ•π‘’ πœ•π‘‘ = 𝑓(𝑒) + 𝐷 πœ•2 𝑒 πœ•π‘₯2 , 0 ≀ π‘₯ ≀ 𝐿, (2)
  • 3.  ISSN: 2088-8708 Int J Elec & Comp Eng, Vol. 10, No. 6, December 2020 : 6260 - 6267 6262 With the following boundary and initial conditions: 𝑒(π‘₯, 𝑑) = 0, 𝑒(𝐿, 𝑑) = 0, 𝑒(π‘₯, 0) = 𝑒₀(π‘₯), where 𝑒 denotes the population density of a species, 𝑓(𝑒) is the species dynamics and 𝐷 is the diffusion coefficient which measure the dispersal efficiency of the relevant species, 𝐿 is the length of the domain. He considered the different boundary conditions which consider the influence of the region exterior to the reaction diffusion domain. He showed that if the domain size is not large enough, then (2) cannot generate spatial patterns and so he found the condition on 𝐿 such that the species will not become extinct if: 𝐿 > πœ‹βˆš 𝐷 𝑓′(0) In this paper, an age-structured model is used to derive a delay advection reaction-diffusion equation with linear advection term. In addition, we find the minimum value of 𝐿 that prevents extinction of the species under the effect of advection term for the reaction-diffusion equation. 2. A DELAY ADVECTION REACTION-DIFFUSION EQUATION WITH LINEAR ADVECTION TERM In this section, we derive a delay advection reaction-diffusion equation with linear advection term from an age-structured model, and then later on, we study and analyze our new model under the effect of the advection term. Define 𝑒 π‘š(π‘₯, 𝑑) to be the intensity of mature adults, with age of at least Ο„. Then it can be represented as follows: 𝑒 π‘š(π‘₯, 𝑑) = ∫ 𝑒(π‘₯, 𝑑, π‘Ž)π‘‘π‘Ž ∞ 𝜏 , (3) where 𝑒(π‘₯, 𝑑, π‘Ž) is the intensity of individuals of age π‘Ž at point π‘₯, time 𝑑 and 𝜏 is the length of the juvenile period. Now, let 𝑒 satisfies the following equation, see Metz [24] and McNair [25]: πœ•π‘’ πœ•π‘‘ + πœ•π‘’ πœ•π‘Ž = π‘ˆ πœ•π‘’ πœ•π‘₯ + 𝑑𝑖 πœ•2 𝑒 πœ•π‘₯2 βˆ’ 𝛾𝑒, 0 < π‘Ž < 𝜏, (4) and that the mature evolve according to: πœ•π‘’ π‘š πœ•π‘‘ = π‘ˆ πœ•π‘’ π‘š πœ•π‘₯ + 𝑑 π‘š πœ•2 𝑒 π‘š πœ•π‘₯2 + 𝑒(π‘₯, 𝑑, 𝜏) βˆ’ 𝛽𝑒 π‘š 2 , π‘₯ ∈ (βˆ’βˆž, ∞), 𝑑 > 0 (5) with 𝑒(π‘₯, 𝑑, 0) = 𝑓(𝑒 π‘š(π‘₯, 𝑑)) (6) the term 𝑒(π‘₯, 𝑑, 𝜏) in (5) represents adult recruitment, since it is those of age 𝜏 (maturation age) and 𝑓(𝑒 π‘š) in (6) refers to the rare of the birth, supposed to depend only on the number of matures at the time. Let 𝑣(π‘₯, π‘Ÿ, π‘Ž) = 𝑒(π‘₯, π‘Ž + π‘Ÿ, π‘Ž), then πœ•π‘£ πœ•π‘Ž = [ πœ•π‘’ πœ•π‘‘ (π‘₯, 𝑑, π‘Ž) + πœ•π‘’ πœ•π‘Ž (π‘₯, 𝑑, π‘Ž)] 𝑑=π‘Ž+π‘Ÿ = π‘ˆ πœ•π‘’ πœ•π‘₯ (π‘₯, π‘Ž + π‘Ÿ, π‘Ž) + 𝑑𝑖 πœ•2 𝑒 πœ•π‘₯2 (π‘₯, π‘Ž + π‘Ÿ, π‘Ž) βˆ’ 𝛾𝑒(π‘₯, π‘Ž + π‘Ÿ, π‘Ž), which implies πœ•π‘£ πœ•π‘Ž = π‘ˆ πœ•π‘£ πœ•π‘₯ + 𝑑𝑖 πœ•2 𝑣 πœ•π‘₯2 βˆ’ 𝛾𝑣 πœ•π‘’ πœ•π‘‘ + πœ•π‘’ πœ•π‘Ž = π‘ˆ πœ•π‘’ πœ•π‘₯ + 𝑑𝑖 πœ•2 𝑒 πœ•π‘₯2 βˆ’ 𝛾𝑒, 0 < π‘Ž < 𝜏, (7)
  • 4. Int J Elec & Comp Eng ISSN: 2088-8708  A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh) 6263 Now, we apply the Fourier transform: 𝑣̂(𝑠, π‘Ÿ, π‘Ž) = ∫ 𝑣(π‘₯, π‘Ÿ, π‘Ž) π‘’βˆ’π‘–π‘ π‘₯ 𝑑π‘₯ ∞ βˆ’βˆž (8) To (8) gives: πœ•π‘£Μ‚ πœ•π‘Ž = (π‘ˆπ‘–π‘  βˆ’ 𝑑𝑖 𝑠² βˆ’ 𝛾)𝑣̂ (9) and 𝑣̂(𝑠, π‘Ÿ, 0) = 𝑒̂(𝑠, π‘Ÿ, 0) = β„±{𝑓(𝑒 π‘š(π‘₯, π‘Ÿ))} (10) The solution of (9) is: 𝑣̂(𝑠, π‘Ÿ, π‘Ž) = β„±{𝑓(𝑒 π‘š(π‘₯, π‘Ÿ))}𝑒(π‘ˆπ‘–π‘ βˆ’π‘‘ 𝑖 π‘ Β²βˆ’π›Ύ)π‘Ž . (11) Now, ℱ⁻¹(𝑒(π‘ˆπ‘–π‘ βˆ’π‘‘ 𝑖 π‘ Β²βˆ’π›Ύ)π‘Ž ) = 1 2πœ‹ 𝑒(π‘ˆπ‘–π‘ βˆ’π‘‘ 𝑖 π‘ Β²βˆ’π›Ύ)π‘Ž 𝑒 𝑖𝑠π‘₯ 𝑑𝑠 = π‘’βˆ’π›Ύπ‘Ž √4πœ‹π‘‘π‘– π‘Ž π‘’βˆ’(π‘₯+π‘ˆπ‘Ž)Β²/(4𝑑 𝑖 π‘Ž) . By using convolution theorem: 𝑣(π‘₯, π‘Ÿ, π‘Ž) = π‘’βˆ’π›Ύπ‘Ž ∫ 1 √4πœ‹π‘‘π‘– π‘Ž ∞ βˆ’βˆž 𝑒 βˆ’(π‘ˆπ‘Ž +π‘₯βˆ’π‘¦)2 4𝑑 𝑖 π‘Ž 𝑓(𝑒 π‘š(𝑦, π‘Ÿ))𝑑𝑦, Therefore 𝑒(π‘₯, 𝑑, 𝜏) = 𝑣(π‘₯, 𝑑 βˆ’ 𝜏, 𝜏) = π‘’βˆ’π›Ύπœ ∫ 1 √4πœ‹π‘‘π‘– 𝜏 ∞ βˆ’βˆž 𝑒 βˆ’(π‘ˆπœ +π‘₯βˆ’π‘¦)2 4𝑑 𝑖 𝜏 𝑓(𝑒 π‘š(𝑦, 𝑑 βˆ’ 𝜏))𝑑𝑦. (12) If 𝑓(𝑒 π‘š) = 𝛼𝑒 π‘š, then the birth rate is symmetric to the number of the mature adults given by 𝛼𝑒 π‘š. Most of the populations, such presumption is valid only when population numbers are relatively little or when there is abundance of nourishment. Inserting (12) with 𝑓(𝑒 π‘š) = 𝛼𝑒 π‘š into (4) yields: πœ•π‘’ π‘š πœ•π‘‘ = π‘ˆ πœ•π‘’ π‘š πœ•π‘₯ + 𝑑 π‘š πœ•2 𝑒 π‘š πœ•π‘₯2 βˆ’ 𝛽𝑒 π‘š 2 +∝ π‘’βˆ’π›Ύπœ ∫ 1 √4πœ‹π‘‘π‘– 𝜏 ∞ βˆ’βˆž 𝑒 βˆ’(π‘ˆπœ +π‘₯βˆ’π‘¦)2 4𝑑 𝑖 𝜏 𝑒 π‘š(𝑦, 𝑑 βˆ’ 𝜏)𝑑𝑦 (13) By applying the following substitution in (13): π‘ˆπœ + π‘₯ βˆ’ 𝑦 = π‘€βˆš4𝑑𝑖 𝜏 , we obtain: π›Όπ‘’βˆ’π›Ύπœ ∫ 1 √4πœ‹π‘‘π‘– 𝜏 ∞ βˆ’βˆž 𝑒 βˆ’(π‘ˆπœ +π‘₯βˆ’π‘¦)2 4𝑑 𝑖 𝜏 𝑒 π‘š(𝑦, 𝑑 βˆ’ 𝜏)𝑑𝑦 = π›Όπ‘’βˆ’π›Ύπœ ∫ 1 √ πœ‹ ∞ βˆ’βˆž π‘’βˆ’π‘€2 𝑒 π‘š(π‘ˆπœ + π‘₯ βˆ’ π‘€βˆš4𝑑𝑖 𝜏, 𝑑 βˆ’ 𝜏)𝑑𝑀. (14) This form of the integral allows us to make approximation for small values of the parameters. Let us approximate (14) for small values of the immature diffusivity 𝑑𝑖:
  • 5.  ISSN: 2088-8708 Int J Elec & Comp Eng, Vol. 10, No. 6, December 2020 : 6260 - 6267 6264 𝑒 π‘š(π‘₯ + π‘ˆπœ βˆ’ π‘€βˆš4𝑑𝑖 𝜏, 𝑑 βˆ’ 𝜏) = 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) βˆ’ π‘€βˆš4𝑑𝑖 𝜏 πœ•π‘’ π‘š πœ•π‘₯ (π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) + 2𝑑𝑖 πœπ‘€2 πœ•2 𝑒 π‘š πœ•π‘₯2 (π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) Therefore, the integral term in (14) can be written as follows: π›Όπ‘’βˆ’π›Ύπœ ∫ 1 √ πœ‹ ∞ βˆ’βˆž π‘’βˆ’π‘€2 (𝑒 π‘š(π‘₯ + π‘ˆπœ βˆ’ π‘€βˆš4𝑑𝑖 𝜏, 𝑑 βˆ’ 𝜏)) 𝑑𝑀 = π›Όπ‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘ˆπœ + π‘₯, 𝑑 βˆ’ 𝜏) + π›Όπ‘’βˆ’π›Ύπœ 𝑑𝑖 𝜏 πœ•2 𝑒 π‘š πœ•π‘₯2 (π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏)), (15) where we have used the following fact: ∫ π‘€Β²π‘’βˆ’π‘€2 𝑑𝑀 = 1 2 √ πœ‹ ∞ βˆ’βˆž . In the limiting case 𝑑𝑖 β†’ 0 (i.e., immatures not diffusing at all), (13) becomes: πœ•π‘’ π‘š πœ•π‘‘ = π‘ˆ πœ•π‘’ π‘š πœ•π‘₯ + 𝑑 π‘š πœ•2 𝑒 π‘š πœ•π‘₯2 + π‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) βˆ’ 𝛽𝑒2 π‘š (16) In this case 𝑑𝑖 = 0, the term π›Όπ‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) can be understood ecologically as follows: Setting 𝑑𝑖 = 0 means the immature are not diffusing. They are, however, still subject to advection. During the period of immaturity, which is of duration 𝜏, each immature will have moved a distance equal to π‘ˆπœ. It follows that an individual that becomes mature at location x must have been born at location π‘₯ + π‘ˆπœ (since the advection is from right to left). The term: π›Όπ‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏), which is the rate at time 𝑑, position π‘₯, at which individuals become mature, is given simply by the birth rate 𝜏 time units ago at location π‘₯ + π‘ˆπœ, 𝛼𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏), corrected for juvenile mortality which explains the π‘’βˆ’π›Ύπœ factor. If the species is close to extinction and the immature are not diffusing, then (16) becomes: πœ•π‘’ π‘š πœ•π‘‘ = π‘ˆ πœ•π‘’ π‘š πœ•π‘₯ + 𝑑 π‘š πœ•2 𝑒 π‘š πœ•π‘₯2 + π›Όπ‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏), 0 < π‘₯ < 𝐿, 𝑑 > 0. (17) 3. EFFECT OF ADVECTION TERM ON THE DERIVED DELAY ADVECTION REACTION- DIFFUSION EQUATION Starting by studying (17) on homogeneous Dirichlet boundary conditions: 𝑒 π‘š(0, 𝑑) = 0, 𝑒 π‘š(𝐿, 𝑑) = 0, and the initial condition 𝑒 π‘š(π‘₯, 0) = 𝑒 π‘š 0 (π‘₯), π‘₯ ∈ [βˆ’πœ, 0] with 𝑒 π‘š 0 (0) > 0. As a starting point, we shall also consider the case when the juveniles are not subject to advection, in this case, the term 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) of (16) becomes: π›Όπ‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘₯, 𝑑 βˆ’ 𝜏). However, the adults will still be subject to advection. Seeking trial solutions for (16) of the form 𝑒 π‘š(π‘₯, 𝑑) = 𝑒στ πœ™(π‘₯), yields: 𝑑 π‘š πœ™β€²β€²(π‘₯) + π‘ˆ πœ™β€²βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎)πœ™(π‘₯) = 0 (18) The characteristic equation for (18) is 𝑑 π‘š 𝑀² + π‘ˆ 𝑀 + (π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) = 0 (19) and the roots are given by: 𝑀₁, 𝑀₂ = 1 2𝑑 π‘š (βˆ’π‘ˆ Β± (βˆšπ‘ˆ2 βˆ’ 4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) ) Thus, we have two cases for the roots: If 𝑀₁ and 𝑀₂ are real, then the solution for πœ™ satisfying the boundary condition πœ™(0) = 0 will be in the form πœ™(π‘₯) = 𝐴(𝑒 𝑀₁π‘₯ βˆ’ 𝑒 𝑀2 π‘₯ )
  • 6. Int J Elec & Comp Eng ISSN: 2088-8708  A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh) 6265 and in order to satisfy πœ™(𝐿) = 0, we will have the zero solution. If M₁ and Mβ‚‚ are complex, then the solution for πœ™(π‘₯) satisfying the boundary condition πœ™(0) = 0 will be of the form πœ™(π‘₯) = 𝑒 ( βˆ’π‘ˆπ‘₯ 2𝑑 π‘š ) 𝐡 𝑠𝑖𝑛(√ 4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) βˆ’ π‘ˆΒ² 2𝑑 π‘š π‘₯). (20) In order to be able to satisfy πœ™(𝐿) = 0, we need: √(4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) βˆ’ π‘ˆΒ²) 2𝑑 π‘š = π‘›πœ‹ 𝐿 , 𝑛 = 1, 2, 3, …. That is: 4𝑑 π‘š(π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ 𝜎) βˆ’ π‘ˆ2 = 4𝑑 π‘š 2 𝑛2 πœ‹2 𝐿2 , 𝑛 = 1, 2, 3, …, which gives the following equation to be solved for 𝜎: 𝜎 = π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπœ βˆ’ π‘ˆΒ² 4𝑑 π‘š βˆ’ 𝑑 π‘š π‘›Β²πœ‹Β² 𝐿² (21) The solution 𝑒 π‘š(π‘₯, 𝑑) of the linearized (17) is of the following form 𝑒 π‘š(π‘₯, 𝑑) = 𝑒 πœŽπ‘‘ 𝐡𝑛 𝑒 βˆ’π‘ˆπ‘₯ 2𝑑 π‘š 𝑠𝑖𝑛 ( 𝑛 πœ‹π‘₯ 𝐿 ) , 𝑛 = 1, 2, 3, …, where 𝜎 satisfies (20). Now, we are going to find the solution of (17) satisfying the initial data by summing over 𝑛 to obtain: 𝑒 π‘š(π‘₯, 𝑑) = βˆ‘ 𝐡𝑛 𝑒 𝜎 𝑛 𝑑 ∞ 𝑛=1 𝑒 βˆ’π‘ˆπ‘₯ 2𝑑 π‘š 𝑠𝑖𝑛 ( 𝑛 πœ‹π‘₯ 𝐿 ), (22) where 𝜎 𝑛 satisfies (21) and using the initial condition and Fourier coefficients, then the value of 𝐡𝑛 is given by: 𝐡{𝑛} = 2 𝐿 ∫ 𝑒 π‘š 0 (π‘₯) 𝑒 π‘ˆπ‘₯ 2𝑑 π‘š 𝑠𝑖𝑛 ( 𝑛 πœ‹π‘₯ 𝐿 ) 𝑑π‘₯ 𝐿 0 , 𝑛 = 1,2,3, … (23) Substituting this value in (22) yields the final solution of the linearized problem: 𝑒 π‘š(π‘₯, 𝑑) = βˆ‘ ( 2 𝐿 ∫ 𝑒 π‘š 0 (π‘₯) 𝑒 π‘ˆπ‘₯ 2𝑑 π‘š 𝑠𝑖𝑛 ( 𝑛 πœ‹π‘₯ 𝐿 ) 𝑑π‘₯ Γ— 𝐿 0 ∞ 𝑛=1 𝑒 𝜎 𝑛 𝑑 𝑒 βˆ’π‘ˆπ‘₯ 2𝑑 π‘š 𝑠𝑖𝑛 ( 𝑛 πœ‹π‘₯ 𝐿 ) (24) The population will decay to zero if π‘…π‘’πœŽ 𝑛 < 0, for all 𝑛 = 1,2,3, …, where 𝜎 𝑛 is any root of (21). In other words, 𝜎 𝑛satisfies: 𝜎 𝑛 = π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽ 𝑛 𝑑 βˆ’ 𝑑 π‘š 𝑛2 πœ‹2 𝐿2 βˆ’ π‘ˆΒ² 4𝑑 π‘š In order to detect a possible change of linear stability of the zero solution of (17) we set 𝜎 = π‘–πœ”, which gives: π‘ˆ2 4𝑑 π‘š + 𝑑 π‘š 𝑛2 πœ‹2 𝐿2 + 𝑖𝑀 = π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’π‘–π‘€π‘‘ ,
  • 7.  ISSN: 2088-8708 Int J Elec & Comp Eng, Vol. 10, No. 6, December 2020 : 6260 - 6267 6266 and the complex conjugate of above equation is: π‘ˆ2 4𝑑 π‘š + 𝑑 π‘š 𝑛2 πœ‹2 𝐿2 βˆ’ 𝑖𝑀 = π›Όπ‘’βˆ’π›Ύπœ 𝑒 𝑖𝑀𝑑 , Thus, we are eliminating π‘’βˆ’π‘–πœ”πœ from the last two equations, yields: πœ”Β² = π›ΌΒ²π‘’βˆ’2π›Ύπœ βˆ’ [ π‘ˆ2 4𝑑 π‘š + 𝑑 π‘š 𝑛2 πœ‹2 𝐿2 ]Β². To find the conditions on the parameters which ensure that: πœ”Β² < 0, since this means that roots of (21) of the form 𝜎 = π‘–πœ”, with πœ” real, cannot exist. Note that: If πœ”Β² < 0 when n=1 then automatically πœ”Β² < 0 for n=2, 3, 4, …, therefore it is sufficient to set 𝑛 = 1. When 𝑛 = 1, πœ”Β² < 0 and if the following inequality holds: π›Όπ‘’βˆ’π›Ύπœ < π‘ˆ2 4𝑑 π‘š + 𝑑 π‘š πœ‹2 𝐿2 then 𝑒 π‘š(π‘₯, 𝑑) β†’ 0 as 𝑑 β†’ 0 , thus the population will become extinct. So, the species will become extinct in this case if: π‘ˆ (advection speed) is large 𝑑 π‘š (mature diffusion) is either very small or very large 𝐿 (length of domain) is small 𝛼 (birth rate) is too small 𝛾 (juvenile mortality) is large 𝜏 (length of juvenile period) is large. Also, if the following inequality holds: π›Όπ‘’βˆ’π›Ύπœ > π‘ˆ2 4𝑑 π‘š + 𝑑 π‘š πœ‹2 𝐿2 , then 𝜎 𝑛 = π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽ 𝑛 𝑑 βˆ’ 𝑑 π‘š 𝑛2 πœ‹2 𝐿2 βˆ’ π‘ˆΒ² 4𝑑 π‘š with 𝑛 = 1 has a real positive root 𝜎. This can be shown by drawing the graphs of π‘ˆ2 4𝑑 π‘š + 𝑑 π‘š πœ‹2 𝐿2 + 𝜎 and π›Όπ‘’βˆ’π›Ύπœ π‘’βˆ’πœŽπ‘‘ against 𝜎, where we can see that the two functions have a real positive root Οƒ which means that the zero solution of (17) is unstable. So, we can conclude that the population will survive in this case if π‘ˆ (advection speed) is too small. 𝐿 (domain) is too large. 𝛼 (birth rate) is too large. 4. DISCUSSIONS AND CONCLUSION We have derived a delay advection reaction-diffusion equation with linear advection term from an age-structured model which is given by: πœ•π‘’ π‘š πœ•π‘‘ = π‘ˆ πœ•π‘’ π‘š πœ•π‘₯ + 𝑑 π‘š πœ•2 𝑒 π‘š πœ•π‘₯2 + π‘’βˆ’π›Ύπœ 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) βˆ’ 𝛽𝑒2 π‘š (25) and our derived equation as in (24) is studied when the species is close to extinction and the immature are not diffusing on homogeneous Dirichlet boundary conditions 𝑒 π‘š(0, 𝑑) = 0, 𝑒 π‘š(𝐿, 𝑑) = 0 and the initial condition 𝑒 π‘š(π‘₯, 0) = 𝑒 π‘š 0 β‰₯ 0 for π‘₯ ∈ [βˆ’πœ, 0]. Also, we considered the case when juveniles are not subject to advection, so the term 𝑒 π‘š(π‘₯ + π‘ˆπœ, 𝑑 βˆ’ 𝜏) in (24) becomes 𝑒 π‘š(π‘₯, 𝑑 βˆ’ 𝜏). However, the adults will still be subject to advection. The conditions on the parameters for the final solution 𝑒 π‘š(π‘₯, 𝑑) for the linearized (25) that prevents extinction of the species under the defect of advection for the reaction-diffusion are founds, where the conclusion can be summarized as follows: ο€­ The species will become extinct if the advection speed if large, mature diffusion is either very small or very large, length of domain is small, juvenile mortality is large and length of juvenile is large. ο€­ The species will survive if advection speed is small, domain is too large and birth rate is too large.
  • 8. Int J Elec & Comp Eng ISSN: 2088-8708  A stage-structured delayed advection reaction-diffusion model for single species (Raed Ali Alkhasawneh) 6267 REFERENCES [1] Britton, N., β€œReaction-Diffusion Equations and Their Applications to Biology,” Academic Press, 1986. [2] Lewis, M., Petrovskii, S., Potts, J., β€œReaction-Diffusion Models: Single Species,” The Mathematics Behind Biological Invasions, pp. 69-105, 2016. [3] Allen, L., β€œPersistence and extinction in single-species reaction-diffusion models,” Bulletin of Mathematical Bioloy, vol. 45, pp. 209-227, 1983. [4] Aiello, W. G. and Freedman, H. I., β€œA time-delay model of single species growth with stage structure,” Math. Biosci., vol. 101, pp. 139-153, 1990. [5] Fisher, R. A., β€œThe wave of advance of advantageous genes,” Annals of Eugenics, vol. 7, pp. 355-369, 1937. [6] Kolmogorov, A. Petrovskii, I. and Piscounov, N., β€œA study of the diffusion equation with increase in the amount of substance, and its application to a biological problem,” In V. M. Tikhomirov, editor, Selected Works of A. N. Kolmogorov I, pages 248C270. Kluwer 1991. Translated by V. M. Volosov from Bull. Moscow Univ., Math. Mech. 1, pp. 1-25, 1937. [7] Fisher R. A., β€œThe genetical theory of natural selection,” Oxford University Press, USA, New Ed edition 2000, variorum edition 1999, 1930. [8] Gurney, W. S. C. and Nisbet, R. M., β€œFluctuation periodicity, generation separation, and the expression of Larval competition,” J. Theoretical Popul. Biol., vol. 28, no. 2, pp. 150-180, 1985. [9] Ablowitz, M. J. and Zeppetella, A., β€œExplicit solutions of Fisher’s equation for a special wave speed,” Bull. Math. Biol., vol. 41, pp. 835-840, 1979. [10] Al-Omari, J. F. M. and Gourley S. A., β€œA nonlocal reaction-diffusion model for a single species with stage structure and distributed maturation delay,” Euro. J. Appl. Math., vol. 16, no. 1, pp. 37-51, 2005. [11] Al-Omari, J., Gourley, S., β€œDynamics of a stage-structured population model incorporating a state-dependent maturation delay,” Nonlinear Anal. Real World Appl., vol. 6, no. 1, pp. 13-33, 2005. [12] Aiello, W. G., Freedman, H. I. and Wu, J., β€œAnalysis of a model representing stage-structured population growth with state-dependent time delay,” SIAM J. Appl. Math., vol. 52, pp. 855-869, 1992. [13] Gourley, S. A., Jennings, R. and Liu, R., β€œAn Advection and Age-Structured Approach to Modeling Bird Migration and Indirect Transmission of Avian Influenza,” SIAM J. Appl. Math., vol. 75, no. 4, pp. 1620-1647, 2005. [14] Sarfaraz W, Madzvamuse A., β€œClassification of parameter spaces for a reaction diffusion model onstationary domains,” Chaos Solitons Fractals, vol. 103, pp. 33-51, 2007. [15] Chunwei W., β€œA stage-structured delayed reaction-diffusion model for A stage-structured delayed reaction- diffusion model for competition between two species,” Electronic Theses and Dissertations, University of Louisville, 2013. [16] Rattanakul, C., Lenbury, Y. and Suksamran, J., β€œAnalysis of advection-diffusion-reaction model for fish population movement with impulsive tagging: stability and traveling wave solution,” Advances in Difference Equations, vol. 2019, 2019. [17] Venkataraman, C., β€œReaction-diffusion system on evolving dmains with applications to the theory of biological pattern formation,” Ph.D. thesis, University of Sussex, 2011. [18] Boonrangsiman, S., Bunwon, K. and Elvin, J., β€œA bifurcation path to chaos in a time-delay fisheries predator-prey model with prey consumption by immature and mature predators,” Math. Computers Simul., vol. 124, pp. 16-29, 2016. [19] Zhang, L. and Zhao, H. Y., β€œTraveling wave solutions in a stage-structured delayed reaction-diffusion model with advection,” J. Math. Modelling Anal., vol. 20, no. 2, pp. 168-187, 2015. [20] Cangiani, A., Georgoulis, E., Morozov, A. and Sutton O., β€œRevealing new dynamical patterns in a reaction- diffusion model with cyclic competition via a novel computational framework,” Proc.R.Soc.A474, 2018. [21] Murray, J. D., β€œMathematical Biology,” 2nd ed., Springer, Berlin, 1993. [22] Murray, J. D., β€œMathematical Biology,” I. An introduction. 3rd ed. Inter, disciplinary Applied Mathematics, Springer, New York, 2002. [23] Murray, J. D. , β€œMathematical biology,” Spatial models and biomedical applications. Springer, New York, 2013. [24] Metz, J. A. J and Diekmann, O., β€œThe Dynamics of Physiologically Structured Populations,” Springer-Verlag, New York, 1986. [25] McNair, J., N., and Goulden, C. E., β€œThe dynamics of age-structured populations with a gestation period: Density- independent growth and egg ration methods for estimating the birth rate,” Theoret. Population Bio., vol. 39, pp. 1-29, 1991.