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Variation in fuel value index of five tree species in Mali: 
               Variation in fuel value index of five tree species in Mali
                          relationships with rainfall gradients
               Carmen Sotelo Montes1, John C. Weber1 , Dimas Agostinho da Silva2, Clarice Andrade2, 
                     Rosilei A. Garcia3, Graciela Inês Bolzón de Muñiz2, Antoine Kalinganire1
                 1World Agroforestry            Centre (Bamako, Mali), 2Universidade Federal do Paraná (Curitiba, Brazil), 
                                         3Universidade Federal Rural do Rio de Janeiro (Rio de Janeiro, Brazil) 

                                          Introduction
Rural communities in the West African Sahel depend on many native tree species for
fuelwood. Tree growth and wood properties such as density, moisture and ash content have
an effect on the fuel’s value. Rainfall varies with latitude and longitude in the region, so one
would expect that tree growth and wood properties of trees in natural populations would
also vary with latitude and longitude. Understanding variation among natural populations is
important for tree domestication and conservation programmes, but there has been very
little research in the West African Sahel. ICRAF initiated a project to investigate geographic
variation in growth and wood properties of native tree species in the Sahel. In this poster,
we present preliminary results of a study of variation in the fuel value index of five native
                                                                                                           B. aegyptiaca
tree species in Mali. The major objective was to determine if the fuel value index of the                                                                                                                                                                                                        P. reticulatum
wood varied with rainfall gradients. Five species were studied: Balanites aegyptiaca (L.)
Delile, Combretum glutinosum Perr.ex DC., Guiera senegalensis J.F. Gmel., Piliostigma
reticulatum (DC.) Hochst. and Ziziphus mauritiana Lam. All five species are used for
firewood, produce abundant natural regeneration if protected from animal browsing and
resprout vigorously after coppicing. Therefore, there is potential to manage natural
regeneration of the species for fuelwood production in rural communities.
                                                                                                         C. glutinosum

                                   Materials and Methods
Trees were sampled in natural populations (i.e. parkland agroforests and woodlands) in five
regions in Mali (Figure 1). In general, sampling was done along a latitudinal transect in each
                                                                                                                                                                                                                                                                                                 Z. mauritiana
region: 15‐16 trees per species per region. Trees were selected if the stem diameter was 4‐
12 cm at 30 cm above ground, was not d
                   b           d           damaged and was not a resprout. A sample of the
                                                  d d                                l f h
wood was obtained from the stem (30‐40 cm above ground) for determination of wood
properties. For each tree, latitude, longitude and elevation were recorded with a GPS                                                              Figure 1. Geographic location of five tree species sampled in five regions in 
receiver, and estimated mean annual rainfall was obtained from the WorldClim database.                   G. senegalensis                           Mali, and mean annual rainfall isohyets (mm) in the sample region. 
Based on Pearson correlation coefficients between geographical coordinates and estimated
                                                                                                     Table 1. Mean values of wood properties of five tree species sampled in                                 Table 2. Regression equations predicting fuel value index of four tree species from
mean annual rainfall at the location of sampled trees, mean annual rainfall decreases with           Mali.                                                                                                   geographical coordinates and mean annual rainfall.
latitude and elevation and increases with longitude (r = ‐0.67, ‐0.12 and 0.41, respectively;        Species         FVI      BDen (kg  NetCV (MJ  AshC            MoistC
                                                                                                                                                                                                             Species                  Equation                                          R2       P         SE
P<0.001 for latitude and longitude and 0.016 for elevation, N=395).                                                           m‐3)       kg‐1)          (%)        (%)
                                                                                                                                                                                                             B. aegyptiaca            1208.85709 – 4.57474(Lon2)                        0.238    <0.001    0.93417
                                                                                                     B. aegyptiaca               881         615            17.7                1.2          11.3            C. glutinosum            859.28441 + 0.00002108(Lat2Elev2)                 0.119    0.002     0.00000655
The fuel value index (FVI) = [(BDen)*(NetCV)]/[(AshC)(MoistC)] where BDen = basic density            C. glutinosum               1191        666            17.7                1.1          10.8            G. senegalensis          1841.45073 + 0.00199(ElevRain)                    0.057    0.034     0.00091861
(kg m‐3), NetCV = net calorific value (MJ kg‐1), AshC = ash content (%) and MoistC = moisture        G. senegalensis 2312                    674            18.6                0.5          11.0
                                                                                                                                                                                                             P. reticulatum           728.21833 – 2.28136(LatLon)                       0.157    <0.001    0.60291
content (%) of the wood. BDen was measured on a disk (2 cm thick, sampled at 30 cm above             P. reticulatum              472         531            17.7                1.9          11.7
                                                                                                                                                                                                             Independent variables: Lat and Lat2 = linear and quadratic latitude (°N), Lon and Lon2 =
ground) using the water‐displacement method. NetCV, AshC and MoistC were measured on                 Z. mauritiana               907         582            18.1                1.2          10.9            linear and quadratic longitude (°W), Elev and Elev2 = linear and quadratic elevation (m),
sawdust (sampled 32‐40 cm above ground). NetCV is the energy available from an air‐dry               FVI = fuel value index BDen = basic density NetCV = net calorific value
                                                                                                                      index,             density,                      value,
                                                                                                                                                                                                             Rain = estimated mean annual rainfall (mm). R2 = coefficient of determination of model.
sample: it was measured using an adiabatic bomb calorimeter. AshC and MoistC were                    AshC = ash content, MoistC = moisture content. Sample size: 79 for all
                                                                                                                                                                                                             P = probability of F ratio. SE = standard error of regression coefficient. Sample size: 79 for
                                                                                                                                                                                                             all species.
measured using an incinerator. Tree age was estimated from the number of rings on the disk           species.
(at 30 cm above ground): estimated mean age of all trees was 5.9 years. Trees varied in age
so the effect of age on the wood variables was investigated using analysis of covariance                                                Balanites aegyptica                                                                                             Combretum glutinosum
                                                                                                                                                                                                                      1600
(ANCOVA) within each region. If the effect was significant (P<0.05), the variable was                    1200
                                                                                                                                                                                                                      1400
adjusted using the covariate parameter from the ANCOVA (yadj = y – βregion(tree age – mean
                                                                                                         1000
age across regions). After adjusting the variables, geographic variation was investigated                                                                                                                             1200

using multiple regression analysis. Regression models were determined using the stepwise                  800                                                                                                         1000

procedure: independent variables included linear, quadratic and interaction terms for the                                                                                                                                800
                                                                                                   FVI




                                                                                                          600
trees’ latitude, longitude, elevation and estimated rainfall.
                                                                                                                                                                                                                FVI




                                                                                                                                                                                                                         600
                                                                                                          400
                                                                                                                                                                                                                         400
                                                                                                                                                                                                                                                                                                     150 m
                                  Results and Discussion                                                  200                                                                                                            200
                                                                                                                                                                                                                                                                                                     350 m
Mean FVI was highest for G. senegalensis and lowest for P. reticulatum (Table 1) due to the                 0                                                                                                             0
large differences in BDen NetCV and AshC between these species Due to rounding error
                     BDen,                                  species.                  error,                        0        2           4           6         8           10         12            14                         11.5
                                                                                                                                                                                                                               11 5    12        12.5
                                                                                                                                                                                                                                                 12 5           13        13.5
                                                                                                                                                                                                                                                                          13 5     14        14.5
                                                                                                                                                                                                                                                                                             14 5
mean FVI in the table does not equal the product of BDen, NetCV, AshC and MoistC.                                                                   Longitude (°W)                                                                                             Latitude (°N)

                                                                                                                                                                                                                                                         Piliostigma reticulatum
FVI of trees was significantly related to latitude, longitude, elevation and/or mean annual                                                    Guiera senegalensis                                                       700
rainfall for four of the five species (Table 2). Judging from the R2, the strongest relationship         3000

                                                                                                                                                                                                                         600
was for B. aegyptiaca while the weakest relationship was for G. senegalensis. Plots of the               2500

regression equations (Figure 2) indicate that FVI did not vary in a consistent manner with                                                                                                                               500

geographical coordinates and rainfall in all four species. The FVI of B. aegyptiaca and C.               2000
                                                                                                                                                                                                                         400
                                                                                                                                                                                                                   FVI




glutinosum was greater in the drier parts of the sample region (i.e. in the east and north):
                                                                                                   FVI




                                                                                                         1500
                                                                                                                                                                                                                         300
FVI of B. aegyptiaca increased from west to east, and FVI of C. glutinosum increased from
south to north especially at higher elevations. The FVI of P. reticulatum also increased from            1000                                                                                             600 mm         200
                                                                                                                                                                                                                                                                                                                 5°W
west to east, but decreased slightly from south to north. In contrast, the FVI of G.                                                                                                                      1000 mm                                                                                                12°W
                                                                                                          500                                                                                                            100
senegalensis was slightly greater in areas of higher rainfall, but the regression equation                                                                                                                                 0
explained very little variation among trees (Table 2). Moreover, there was no significant                  0
                                                                                                                0       50        100        150     200    250      300        350    400          450                        11.5         12          12.5         13          13.5           14        14.5
relationship f Z mauritiana.
  l ti hi for Z.          iti                                                                                                                      Elevation (m)                                                                                                Latitude  (°N)

                                                                                                         Figure 2. Fuel value index (FVI) of four tree species predicted by geographical coordinates and mean annual rainfall in Mali
                                        Conclusion                                                       (regression equations in Table 2).
We cannot make a general statement about the relationship between FVI of trees of these
species and rainfall gradients in Mali. Relationships depend on the specific species and the
variation in wood properties that affect FVI in that species. The species with higher FVI are
of course exploited more for charcoal and firewood production. Therefore, researchers
should work with rural communities to develop participatory domestication/conservation
strategies tailored for each species and sustainable fuelwood production systems for species
                                                                                                                                               Research funded by a post‐doctoral research grant from the World Agroforestry Centre,
with higher FVI.                                                                                                                               the International Fund for Agricultural Development, UFPR and UFRRJ. Contact
                                                                                                                                               c.sotelo@cgiar.org for further details.

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  • 1. Variation in fuel value index of five tree species in Mali:  Variation in fuel value index of five tree species in Mali relationships with rainfall gradients Carmen Sotelo Montes1, John C. Weber1 , Dimas Agostinho da Silva2, Clarice Andrade2,  Rosilei A. Garcia3, Graciela Inês Bolzón de Muñiz2, Antoine Kalinganire1 1World Agroforestry Centre (Bamako, Mali), 2Universidade Federal do Paraná (Curitiba, Brazil),  3Universidade Federal Rural do Rio de Janeiro (Rio de Janeiro, Brazil)  Introduction Rural communities in the West African Sahel depend on many native tree species for fuelwood. Tree growth and wood properties such as density, moisture and ash content have an effect on the fuel’s value. Rainfall varies with latitude and longitude in the region, so one would expect that tree growth and wood properties of trees in natural populations would also vary with latitude and longitude. Understanding variation among natural populations is important for tree domestication and conservation programmes, but there has been very little research in the West African Sahel. ICRAF initiated a project to investigate geographic variation in growth and wood properties of native tree species in the Sahel. In this poster, we present preliminary results of a study of variation in the fuel value index of five native B. aegyptiaca tree species in Mali. The major objective was to determine if the fuel value index of the P. reticulatum wood varied with rainfall gradients. Five species were studied: Balanites aegyptiaca (L.) Delile, Combretum glutinosum Perr.ex DC., Guiera senegalensis J.F. Gmel., Piliostigma reticulatum (DC.) Hochst. and Ziziphus mauritiana Lam. All five species are used for firewood, produce abundant natural regeneration if protected from animal browsing and resprout vigorously after coppicing. Therefore, there is potential to manage natural regeneration of the species for fuelwood production in rural communities. C. glutinosum Materials and Methods Trees were sampled in natural populations (i.e. parkland agroforests and woodlands) in five regions in Mali (Figure 1). In general, sampling was done along a latitudinal transect in each Z. mauritiana region: 15‐16 trees per species per region. Trees were selected if the stem diameter was 4‐ 12 cm at 30 cm above ground, was not d b d damaged and was not a resprout. A sample of the d d l f h wood was obtained from the stem (30‐40 cm above ground) for determination of wood properties. For each tree, latitude, longitude and elevation were recorded with a GPS Figure 1. Geographic location of five tree species sampled in five regions in  receiver, and estimated mean annual rainfall was obtained from the WorldClim database. G. senegalensis Mali, and mean annual rainfall isohyets (mm) in the sample region.  Based on Pearson correlation coefficients between geographical coordinates and estimated Table 1. Mean values of wood properties of five tree species sampled in Table 2. Regression equations predicting fuel value index of four tree species from mean annual rainfall at the location of sampled trees, mean annual rainfall decreases with Mali. geographical coordinates and mean annual rainfall. latitude and elevation and increases with longitude (r = ‐0.67, ‐0.12 and 0.41, respectively; Species FVI BDen (kg  NetCV (MJ  AshC  MoistC Species Equation R2 P SE P<0.001 for latitude and longitude and 0.016 for elevation, N=395). m‐3) kg‐1) (%) (%) B. aegyptiaca 1208.85709 – 4.57474(Lon2) 0.238 <0.001 0.93417 B. aegyptiaca 881 615 17.7 1.2 11.3 C. glutinosum 859.28441 + 0.00002108(Lat2Elev2) 0.119 0.002 0.00000655 The fuel value index (FVI) = [(BDen)*(NetCV)]/[(AshC)(MoistC)] where BDen = basic density C. glutinosum 1191 666 17.7 1.1 10.8 G. senegalensis 1841.45073 + 0.00199(ElevRain) 0.057 0.034 0.00091861 (kg m‐3), NetCV = net calorific value (MJ kg‐1), AshC = ash content (%) and MoistC = moisture G. senegalensis 2312 674 18.6 0.5 11.0 P. reticulatum 728.21833 – 2.28136(LatLon) 0.157 <0.001 0.60291 content (%) of the wood. BDen was measured on a disk (2 cm thick, sampled at 30 cm above P. reticulatum 472 531 17.7 1.9 11.7 Independent variables: Lat and Lat2 = linear and quadratic latitude (°N), Lon and Lon2 = ground) using the water‐displacement method. NetCV, AshC and MoistC were measured on Z. mauritiana 907 582 18.1 1.2 10.9 linear and quadratic longitude (°W), Elev and Elev2 = linear and quadratic elevation (m), sawdust (sampled 32‐40 cm above ground). NetCV is the energy available from an air‐dry FVI = fuel value index BDen = basic density NetCV = net calorific value index, density, value, Rain = estimated mean annual rainfall (mm). R2 = coefficient of determination of model. sample: it was measured using an adiabatic bomb calorimeter. AshC and MoistC were AshC = ash content, MoistC = moisture content. Sample size: 79 for all P = probability of F ratio. SE = standard error of regression coefficient. Sample size: 79 for all species. measured using an incinerator. Tree age was estimated from the number of rings on the disk species. (at 30 cm above ground): estimated mean age of all trees was 5.9 years. Trees varied in age so the effect of age on the wood variables was investigated using analysis of covariance Balanites aegyptica Combretum glutinosum 1600 (ANCOVA) within each region. If the effect was significant (P<0.05), the variable was 1200 1400 adjusted using the covariate parameter from the ANCOVA (yadj = y – βregion(tree age – mean 1000 age across regions). After adjusting the variables, geographic variation was investigated 1200 using multiple regression analysis. Regression models were determined using the stepwise 800 1000 procedure: independent variables included linear, quadratic and interaction terms for the 800 FVI 600 trees’ latitude, longitude, elevation and estimated rainfall. FVI 600 400 400 150 m Results and Discussion 200 200 350 m Mean FVI was highest for G. senegalensis and lowest for P. reticulatum (Table 1) due to the 0 0 large differences in BDen NetCV and AshC between these species Due to rounding error BDen, species. error, 0 2 4 6 8 10 12 14 11.5 11 5 12 12.5 12 5 13 13.5 13 5 14 14.5 14 5 mean FVI in the table does not equal the product of BDen, NetCV, AshC and MoistC. Longitude (°W) Latitude (°N) Piliostigma reticulatum FVI of trees was significantly related to latitude, longitude, elevation and/or mean annual Guiera senegalensis 700 rainfall for four of the five species (Table 2). Judging from the R2, the strongest relationship 3000 600 was for B. aegyptiaca while the weakest relationship was for G. senegalensis. Plots of the 2500 regression equations (Figure 2) indicate that FVI did not vary in a consistent manner with 500 geographical coordinates and rainfall in all four species. The FVI of B. aegyptiaca and C. 2000 400 FVI glutinosum was greater in the drier parts of the sample region (i.e. in the east and north): FVI 1500 300 FVI of B. aegyptiaca increased from west to east, and FVI of C. glutinosum increased from south to north especially at higher elevations. The FVI of P. reticulatum also increased from 1000 600 mm 200 5°W west to east, but decreased slightly from south to north. In contrast, the FVI of G. 1000 mm 12°W 500 100 senegalensis was slightly greater in areas of higher rainfall, but the regression equation 0 explained very little variation among trees (Table 2). Moreover, there was no significant 0 0 50 100 150 200 250 300 350 400 450 11.5 12 12.5 13 13.5 14 14.5 relationship f Z mauritiana. l ti hi for Z. iti Elevation (m) Latitude  (°N) Figure 2. Fuel value index (FVI) of four tree species predicted by geographical coordinates and mean annual rainfall in Mali Conclusion (regression equations in Table 2). We cannot make a general statement about the relationship between FVI of trees of these species and rainfall gradients in Mali. Relationships depend on the specific species and the variation in wood properties that affect FVI in that species. The species with higher FVI are of course exploited more for charcoal and firewood production. Therefore, researchers should work with rural communities to develop participatory domestication/conservation strategies tailored for each species and sustainable fuelwood production systems for species Research funded by a post‐doctoral research grant from the World Agroforestry Centre, with higher FVI. the International Fund for Agricultural Development, UFPR and UFRRJ. Contact c.sotelo@cgiar.org for further details.